Physcia clementei (Turner) Lynge

in Rabenhorst Kryptogamenfl., ed. 2: 93, 1935. Basionym: Parmelia clementei Turner in Smith & Sowerby - Engl. Bot., 25: tab. 1779, 1807.
Synonyms: Hagenia stellaris var. caricae Schaer. ex A. Massal.; Parmelia sideralis Ach.; Physcia astroidea auct.; Physcia clementiana (Ach.) J. Kickx f.
Distribution: N - VG (Castello 1996, Castello & Skert 2005), Frl (Badin & Nimis 1996), Ven (Lazzarin 2000, Nascimbene & Marini 2010), Lomb (Zocchi & al. 1997, Arosio & al. 2003), Piem (Arosio & al. 1998, Castino 2004, Morisi 2005, Furlanetto 2010), Emil (Nimis & al. 1996, Gerdol & al. 2014, Fariselli & al. 2020), Lig (Giordani & al. 2001, 2002, Brunialti & Giordani 2003, Giordani 2006, Giordani & Incerti 2008). C - Tosc (Loppi & Putortì 1995, Loppi & al. 1995, 1997, 1998b, 2002b, 2002c, Loppi 1996, Loppi & De Dominicis 1996, Bacci & al. 2000, Benesperi 2000a, Lorenzini & al. 2003, Brunialti & Frati 2010, Loppi & Nascimbene 2010, Paoli & al. 2012, 2015d), Marc, Laz (Massari & Ravera 2002, Ravera & al. 2003, Ruisi & al. 2005, Munzi & al. 2007, Gagliardi & al. 2010), Mol (Frati & al. 2004, Caporale & al. 2008, Paoli & al. 2015), Sar (Neuwirth 2018). S - Camp (Nimis & Tretiach 2004, Catalano & al. 2016), Si (Nimis & al. 1994, Ravera & al. 2023b).
Description: Thallus foliose, heteromerous, dorsiventral, narrow-lobed, forming orbicular, closely adpressed, up to 4 cm wide rosettes. Lobes flat to concave, 0.5-0.8(-1.2) mm wide, sparingly branched, dull white, the central parts often covered by isidia-like pustules dissolving into granulose soredia that may fuse and cover the whole thallus. Lower surface white at lobe-tips, darker in central parts, with a few, concolorous, mostly simple rhizines. Upper cortex paraplectenchymatous; medulla white; lower cortex pseudoparenchymatous. Apothecia rather rare, lecanorine, to 2 mm across, with a black, sometimes thinly pruinose disc and an often crenulate, isidiate-sorediate margin. Epithecium brown; hymenium and hypothecium colourless; paraphyses slender, simple or forked in upper part, the apical cells clavate, with a thin, dark brown cap. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, brown, ellipsoid, 15-20(-24) x 7-11 µm, Pachysporaria-type. Pycnidia rare. Conidia bacilliform. Photobiont chlorococcoid. Spot tests: opper cortex and medulla K+ yellow, C-, KC-, P+ faintly yellow. Chemistry: uppper cortex with atranorin; medulla with matranorin, lecotylin and traces of other triterpenoids.
Note: a Mediterranean to mild-temperate, mainly western species growing on more or less isolated trees; rare throughout the country, but most frequent in Tyrrhenian Italy and in the upper Friulian plain (humid-rainy climate).
Growth form: Foliose, narrow lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: rather rare
Humid mediterranean belt: very rare
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10538)
2001/12/15



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10538)
2001/12/15
apothecium



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10538)
2001/12/15
soralia



Felix Schumm - CC BY-SA 4.0
[20173], Jugoslavia, Dalmatia, peninsula Pelješac, in fundo vallis pone pagum Žuljana, alt. 20 m. Ad corticem Oleae vetustae. Leg. et det. A. Vezda, 17.07.1979, conf. J. Elix. VEZDA: LICHENES SELECTI EXSICCATI NR. 1762.



Felix Schumm - CC BY-SA 4.0
[20173], Jugoslavia, Dalmatia, peninsula Pelješac, in fundo vallis pone pagum Žuljana, alt. 20 m. Ad corticem Oleae vetustae. Leg. et det. A. Vezda, 17.07.1979, conf. J. Elix. VEZDA: LICHENES SELECTI EXSICCATI NR. 1762.



Felix Schumm - CC BY-SA 4.0
[20173], Jugoslavia, Dalmatia, peninsula Pelješac, in fundo vallis pone pagum Žuljana, alt. 20 m. Ad corticem Oleae vetustae. Leg. et det. A. Vezda, 17.07.1979, conf. J. Elix. VEZDA: LICHENES SELECTI EXSICCATI NR. 1762.



Felix Schumm - CC BY-SA 4.0
[20173], Jugoslavia, Dalmatia, peninsula Pelješac, in fundo vallis pone pagum Žuljana, alt. 20 m. Ad corticem Oleae vetustae. Leg. et det. A. Vezda, 17.07.1979, conf. J. Elix. VEZDA: LICHENES SELECTI EXSICCATI NR. 1762.



Felix Schumm - CC BY-SA 4.0
[20173], Jugoslavia, Dalmatia, peninsula Pelješac, in fundo vallis pone pagum Žuljana, alt. 20 m. Ad corticem Oleae vetustae. Leg. et det. A. Vezda, 17.07.1979, conf. J. Elix. VEZDA: LICHENES SELECTI EXSICCATI NR. 1762.



Felix Schumm - CC BY-SA 4.0
[20173], Jugoslavia, Dalmatia, peninsula Pelješac, in fundo vallis pone pagum Žuljana, alt. 20 m. Ad corticem Oleae vetustae. Leg. et det. A. Vezda, 17.07.1979, conf. J. Elix. VEZDA: LICHENES SELECTI EXSICCATI NR. 1762.



Felix Schumm - CC BY-SA 4.0
[20173], Jugoslavia, Dalmatia, peninsula Pelješac, in fundo vallis pone pagum Žuljana, alt. 20 m. Ad corticem Oleae vetustae. Leg. et det. A. Vezda, 17.07.1979, conf. J. Elix. VEZDA: LICHENES SELECTI EXSICCATI NR. 1762.



Felix Schumm - CC BY-SA 4.0
[20173], Jugoslavia, Dalmatia, peninsula Pelješac, in fundo vallis pone pagum Žuljana, alt. 20 m. Ad corticem Oleae vetustae. Leg. et det. A. Vezda, 17.07.1979, conf. J. Elix. VEZDA: LICHENES SELECTI EXSICCATI NR. 1762.
at: atranorin, a: hopane-15 ,22-diol, b: 15 -acetoxyhopane-22-ol


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physcia_clementei.htm
France, session AFL 2015 dans le Lot, Blanzaguet


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physcia_clementei.htm
France, session AFL 2015 dans le Lot, Blanzaguet


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Térénez


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Térénez


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Térénez


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Térénez


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Térénez


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Argol


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Châteaulin


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Plougonven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Plougonven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Plougonven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=469&lang=en
France, Dinéault