Physcia dubia (Hoffm.) Lettau
Hedwigia, 52: 254, 1912. Basionym: Lobaria dubia Hoffm. - Deutschl. Fl.: 156, 1796.
Synonyms: Parmelia caesia var. teretiuscula Ach.; Parmelia pulchella var. dubia (Hoffm.) Schaer.; Physcia caesia var. dubia (Hoffm.) Th. Fr.; Physcia caesitia Nyl.; Physcia dubia var. teretiuscula (Ach.) Clauzade & Cl. Roux; Physcia intermedia Vain.; Physcia lyngei Nádv.; Physcia teretiuscula (Ach.) Lynge; Physcia tribacia auct. p.p. non (Ach.) Nyl.; Physcia wahlenbergii Lynge
Distribution: N - VG, Frl (Tretiach & Hafellner 2000), Ven (Nascimbene & Caniglia 1997, 2003c, Caniglia & al. 1999, Nascimbene 2005c, 2008c, Thor & Nascimbene 2007, Nascimbene & Marini 2007), TAA (Caniglia & al. 2002, Nascimbene 2003, 2006c, 2008b, 2014, Nascimbene & al. 2005, 2006, 2006e, 2007b, 2022, Zarabska & al. 2009, Spitale & Nascimbene 2012, Nimis & al. 2015, Ravera & al. 2023b), Lomb (Rivellini 1994, Grieco & Groppali 1995, Brusoni & al. 1997, Zocchi & al. 1997, Roella 1999, Brusoni & Valcuvia 2000, Arosio & al. 2003, Valcuvia & al. 2003, De Vita & Valcuvia 2004, Anderi & al. 2005, Furlanetto 2010, Gheza 2019, 2019b), Piem (Morisi & Sereno 1995, Arosio & al. 1998, Isocrono & Falletti 1999, Piervittori 2003, Giordani & al. 2003b, Castino 2004, Isocrono & al. 2004, 2005b, 2006, 2007, Griselli & al. 2003, Isocrono & Piervittori 2008, Favero-Longo & al. 2009b, Furlanetto 2010, Lich. Ital. Exs. 45: Isocrono & al. 2020), VA (Piervittori & Isocrono 1997, 1999, Valcuvia 2000, Piervittori & al. 2001, Matteucci & al. 2008, 2008c, 2015c), Emil (Dalle Vedove & al. 2002, Fariselli & al. 2020), Lig (Brunialti & al. 1999, Giordani & Brunialti 2000, Brunialti & Giordani 2003, Giordani & al. 2003b). C - Tosc, Umb (Ravera & al. 2006, Panfili 2007), Laz (Ruisi & al. 2005), Abr (Nimis & Tretiach 1999, Gheza & al. 2021), Mol (Ravera & Genovesi 2012, Genovesi & Ravera 2014), Sar (Rizzi & al. 2011, Cossu & al. 2015, Di Nuzzo & al. 2022). S - Bas (Potenza 2006, Paoli & al. 2006, Potenza & Fascetti 2012), Cal (Puntillo 1996), Si (Campisi & al. 2020).
Description: Thallus foliose, heteromerous, dorsiventral, narrow-lobed, rather loosely adnate, forming irregular, rarely orbicular, up to 3(-5) cm wide rosettes. Lobes 1-2 mm wide, whitish grey to dark grey, often paler at margins, mostly epruinose and emaculate, with lip-shaped soralia at the tips of ascending lobes. Lower surface pale, with mostly simple rhizines. Upper cortex paraplectenchymatous; medulla white; lower cortex prosoplectenchymatous. Apothecia very rare, lecanorine, with a usually sorediate thalline margin.Epithecium brown; hymenium and hypothecium colourless; paraphyses slender, simple or forked in upper part, the apical cells clavate, with a thin, dark brown cap. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, brown, ellipsoid, 16-24(-28) x 6-10 µm, the wall thickened at apex and at septum, Physcia-type. Pycnidia rare, immersed. Conidia subcylindrical, 4-6 x 1 µm. Photobiont chlorococcoid. Spot tests: upper cortex K+ yellow, C-, KC-, P- or P+ faintly yellow; medulla K-, C-, KC-, P- Chemistry: upper cortex with atranorin. Note: a widespread holarctic species with a broad latitudinal and altitudinal range, found on base-rich, nutrient-enriched substrata, both in natural situations and on walls in villages, rarely on dust-impregnated bark at the base of wayside trees, with a wide altitudinal range.
Growth form: Foliose, narrow lobed
Substrata: bark and rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: rather common
Subalpine belt: very common
Oromediterranean belt: rather rare
Montane belt: common
Submediterranean belt: rare
Padanian area: extremely rare
Humid submediterranean belt: rather rare
Humid mediterranean belt: very rare
Dry mediterranean belt: absent
Predictive model
Herbarium samples
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33397)
2001/12/07
soralium
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33397)
2001/12/07
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm - CC BY-SA 4.0
[LIAF_28], Afghanistan, Prov. Kabul, Paghman-Gebirge, W des Ortes Paghman, Chap-Darrah. 68°53’ E, 34°37’ N, 2720 m. An n-exponierten, schattigen Felswänden, zusammen mit Physcia caesia (Hoffm.) Hampe. Leg. D. Podlech (Pod 38/1b), 21.06.1970. LICHENOTHECA AFGHANICA NR. 28
Felix Schumm - CC BY-SA 4.0
[LIAF_28], Afghanistan, Prov. Kabul, Paghman-Gebirge, W des Ortes Paghman, Chap-Darrah. 68°53’ E, 34°37’ N, 2720 m. An n-exponierten, schattigen Felswänden, zusammen mit Physcia caesia (Hoffm.) Hampe. Leg. D. Podlech (Pod 38/1b), 21.06.1970. LICHENOTHECA AFGHANICA NR. 28
Felix Schumm - CC BY-SA 4.0
[LIAF_28], Afghanistan, Prov. Kabul, Paghman-Gebirge, W des Ortes Paghman, Chap-Darrah. 68°53’ E, 34°37’ N, 2720 m. An n-exponierten, schattigen Felswänden, zusammen mit Physcia caesia (Hoffm.) Hampe. Leg. D. Podlech (Pod 38/1b), 21.06.1970. LICHENOTHECA AFGHANICA NR. 28
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Physcia_dubia.htm
France, https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physcia_clementei.htm
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Physcia_dubia.htm
France, https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physcia_clementei.htm
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Physcia_dubia.htm
France, https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physcia_clementei.htm
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Physcia_dubia_teret.htm
France, 5/10/2010 - session AFL 2010 en Corse, Haut-Asco
as morpho. teretiuscula
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Physcia_dubia_teret.htm
France, 5/10/2010 - session AFL 2010 en Corse, Haut-Asco
as morpho. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Chapelle Saint-Sebastian
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Plomodiern
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
Italy, Sardinia
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Growth form: Foliose, narrow lobed
Substrata: bark and rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: rather common
Subalpine belt: very common
Oromediterranean belt: rather rare
Montane belt: common
Submediterranean belt: rare
Padanian area: extremely rare
Humid submediterranean belt: rather rare
Humid mediterranean belt: very rare
Dry mediterranean belt: absent
Predictive model
| Herbarium samples |
Index Fungorum
GBIF
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33397)
2001/12/07
soralium
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33397)
2001/12/07
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm – CC BY-SA 4.0
[16522], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung zum Käppelehof, 48,22853° N, 8,31805° E, 670 m, an Granitfelsen. Leg. et det. Schumm 07.07.2010
Felix Schumm - CC BY-SA 4.0
[LIAF_28], Afghanistan, Prov. Kabul, Paghman-Gebirge, W des Ortes Paghman, Chap-Darrah. 68°53’ E, 34°37’ N, 2720 m. An n-exponierten, schattigen Felswänden, zusammen mit Physcia caesia (Hoffm.) Hampe. Leg. D. Podlech (Pod 38/1b), 21.06.1970. LICHENOTHECA AFGHANICA NR. 28
Felix Schumm - CC BY-SA 4.0
[LIAF_28], Afghanistan, Prov. Kabul, Paghman-Gebirge, W des Ortes Paghman, Chap-Darrah. 68°53’ E, 34°37’ N, 2720 m. An n-exponierten, schattigen Felswänden, zusammen mit Physcia caesia (Hoffm.) Hampe. Leg. D. Podlech (Pod 38/1b), 21.06.1970. LICHENOTHECA AFGHANICA NR. 28
Felix Schumm - CC BY-SA 4.0
[LIAF_28], Afghanistan, Prov. Kabul, Paghman-Gebirge, W des Ortes Paghman, Chap-Darrah. 68°53’ E, 34°37’ N, 2720 m. An n-exponierten, schattigen Felswänden, zusammen mit Physcia caesia (Hoffm.) Hampe. Leg. D. Podlech (Pod 38/1b), 21.06.1970. LICHENOTHECA AFGHANICA NR. 28
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Physcia_dubia.htm
France, https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physcia_clementei.htm
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Physcia_dubia.htm
France, https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physcia_clementei.htm
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Physcia_dubia.htm
France, https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physcia_clementei.htm
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Physcia_dubia_teret.htm
France, 5/10/2010 - session AFL 2010 en Corse, Haut-Asco
as morpho. teretiuscula
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_3/Physcia_dubia_teret.htm
France, 5/10/2010 - session AFL 2010 en Corse, Haut-Asco
as morpho. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Chapelle Saint-Sebastian
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Plomodiern
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
Italy, Sardinia
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=470&lang=en
France, Cap Sizun
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1068&lang=en
France, Saint Lary Soulan 1000 m
var. teretiuscula
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
