Physconia grisea (Lam.) Poelt subsp. grisea
Nova Hedwigia, 9: 30, 1965. Basionym: Lichen griseus Lam. - Encycl. Meth. Bot., 3: 480, 1789.
Synonyms: Hagenia pulverulenta var. pityrea (Ach.) Bagl. & Carestia; Parmelia farrea Ach. non auct. p.p.; Parmelia pityrea (Ach.) Ach.; Physcia grisea (Lam.) Zahlbr.; Physcia grisea var. hillmannii (Lynge) Nádv.; Physcia grisea var. pityrea (Ach.) Flagey; Physcia pityrea (Ach.) Nyl.; Physconia farrea (Ach.) Poelt non sensu Poelt; Poeltonia grisea (Lam.) S.Y. Kondr., Lőkös & Hur
Distribution: N - VG (Carvalho 1997, Castello 2002, Martellos & Castello 2004), Frl (Badin & Nimis 1996, Castello & Skert 2005, Bernini & al. 2010), Ven (Nimis & al. 1996c, Nascimbene & Caniglia 1997, Caniglia & al. 1999, Lazzarin 2000, Valcuvia & al. 2000c, Nascimbene 2008), TAA (Nascimbene & al. 2007b, 2022, Zarabska & al. 2009), Lomb (Arosio & Rinaldi 1995, Valcuvia & Brusoni 1996, Brusoni & al. 1997, Zocchi & al. 1997, Brusoni & Valcuvia 2000, Casarini & al. 2000, Arosio & al. 2000, 2003, Anderi & al. 2005, Valcuvia & Truzzi 2007b, Furlanetto 2010, Gheza & al. 2020, 2023, Favero-Longo & al. 2023), Piem (Caniglia & al. 1992, Arosio & al. 1998, Clerc & al. 1999, Roella 1999, Buzio 2003, Castino & Ropolo 2003, Piervittori 2003, Isocrono & al. 2003, 2004, 2007, Griselli & al. 2003, Castino 2004, Furlanetto 2010, Giordani & Malaspina 2016), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Giordani & al. 2003b, Matteucci & al. 2008c), Emil (Bassi 1995, Nimis & al. 1996, Gasparo & Tretiach 1996, Tretiach 1997, Valcuvia & Savino 2000, Sallese 2003, Marconi & al. 2006, Morselli & Regazzi 2006, Cioffi 2009, Malavasi 2014, Gerdol & al. 2014, Fariselli & al. 2020,Lich. Ital. Exs. 60: Isocrono & al. 2021), Lig (Passauer 1976, Castello & al. 1994, Giordani & al. 2001, 2002, Brunialti & Giordani 2003, Giordani 2006, Giordani & Incerti 2008). C - Tosc (Tretiach & Nimis 1994, Loppi & al. 1995, 1996b, 1997, 1997e, 1998b, 2002, 2002b, 2002c, 2003, 2004, 2006, Monaci & al. 1997, Loppi & Nascimbene 1998, 2010, Putortì & Loppi 1999, Dupla Graec. Lich. 64: Obermayer 1999, Bacci & al. 2000, Del Guasta 2001, Loppi & Corsini 2003, Lorenzini & al. 2003, Loppi & Frati 2004, Frati & al. 2006b, 2007, Paoli & Loppi 2008, Lohtander & al. 2008, Lastrucci & al. 2009, Brunialti & Frati 2010, Benesperi 2011, Paoli & al. 2012, 2012b, 2013, 2015d, Brackel 2015, Frati & Brunialti 2023), Marc (Gasparo & al. 1989, Nimis & Tretiach 1999, Brunialti & al. 2012, Brackel 2015), Umb (Ravera 1998, Ravera & al. 2006, Frati & Brunialti 2006, Brackel 2015, Ravera & Ciotti 2015), Laz (Ravera & al. 1999, Massari & Ravera 2002, Ruisi & al. 2005, Munzi & al. 2007, Ravera 2008b, Ravera & Genovesi 2008, Genovesi & al. 2011, Zucconi & al. 2013), Abr (Recchia & al. 1993, Olivieri & Pacioni 1996, Olivieri & al. 1997, 1997b, Loppi & al. 1999, Nimis & Tretiach 1999, Brackel 2015, Caporale & al. 2016), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Paoli & al. 2011, 2015, Genovesi & Ravera 2014, Brackel 2020, Caporale & Ravera 2020), Sar (Zedda 1995, 2002, 2002b, Loi & al. 2000, Zedda & Sipman 2001, Otte & al. 2002, Rizzi & al. 2011, Kodnik & al. 2011, Giordani & al. 2013, Cossu 2013, Di Nuzzo & al. 2022). S - Camp (Ricciardi & al. 2000, Aprile & al. 2003b, 2011, Nimis & Tretiach 2004, Garofalo & al. 2010, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999), Bas (Bartoli & Puntillo 1998, Nimis & Tretiach 1999, Potenza 2006, Paoli & al. 2006, Potenza & al. 2010, Brackel 2011), Cal (Puntillo & Puntillo 2004, Incerti & Nimis 2006), Si (Ottonello & Salone 1994, Nimis & al. 1996b, Grillo & al. 1996, Ottonello 1996, Grillo 1998, Grillo & Caniglia 2004, 2006, Brackel 2008c,Puglisi & Cataldo 2019).
Description: Thallus foliose, heteromerous, dorsiventral, narrow-lobed, forming regular to irregular 5-8 cm wide rosettes. Lobes (1-)2-3 mm wide, radiating and partly overlapping, markedly pruinose at least at tips, flat to irregularly concave, grey-brown to brown, with primarily marginal, elongate soralia tending to become laminal in older parts, the soredia coarsely granular to almost isidioid. Lower surface whitish in marginal parts, pale brown in central parts, with mostly simple, whitish to pale brown rhizines. Upper cortex paraplectenchymatous; medulla white; lower cortex prosoplectenchymatous, with hyphae oriented parallel to the long axis of lobes. Apothecia rare, lecanorine, 1-3 mm across, with a pruinose disc and a sorediate thalline margin. Epithecium brown; hymenium and hypothecium colourless; paraphyses slender, simple or forked in upper part, the apical cells clavate, with a thin, dark brown cap. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, brown, ellipsoid, (25-)27-34 x (12-)14-17 µm, thickened at septum but not at apices, Physconia-type. Pycnidia frequent, immersed, inconspicuous. Conidia 4-7 x 1-2 µm. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-. Chemistry: medulla with four unidentified lichen substances. Note: a mainly mild-temperate, perhaps holarctic lichen found both on bark (often on basal parts of isolated trees) and on calciferous rocks (especially calcareous sandstone, e.g. on walls); widespread throughout the country with optimum below the montane belt, locally common also in urban areas.
Growth form: Foliose, narrow lobed
Substrata: bark and rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: very common
Padanian area: rather rare
Humid submediterranean belt: very common
Humid mediterranean belt: rather common
Dry mediterranean belt: extremely rare
Predictive model
Herbarium samples
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20462)
2001/12/05
detail of soredia
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20462)
2001/12/05
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10537)
2002/11/04
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli-Venezia Giulia, TS, Basovizza; altitude: 2
2008.02.21
with (in upper part of picture) Physconia distorta
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Slovenia, Lipica/Lipizza; altitude: 2
2008.02.21
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Slovenia, Lipica/Lipizza; altitude: 2
2008.02.21
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Slovenia, Lipica/Lipizza; altitude: 2
2008.02.21
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Slovenia, Lipica/Lipizza; altitude: 2
2008.02.21
Antonio Valero; Owner: Antonio Valero
Spain
2010
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physconia_grisea.htm
France, 19/3/2013 - sur Acer - Mont Dore - Puy-de-Dôme
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physconia_grisea.htm
France, 19/3/2013 - sur Acer - Mont Dore - Puy-de-Dôme
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physconia_grisea.htm
France, 19/3/2013 - sur Acer - Mont Dore - Puy-de-Dôme
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physconia_grisea.htm
France, 19/3/2013 - sur Acer - Mont Dore - Puy-de-Dôme
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: https://italic.units.it/flora/?procedure=img_ins2&species_id=2881
France, Quimper
Bernard Bouffinier - Source: https://italic.units.it/flora/?procedure=img_ins2&species_id=2881
France, Quimper
Bernard Bouffinier - Source: https://italic.units.it/flora/?procedure=img_ins2&species_id=2881
France, Quimper
Jacques Haine - Source: https://italic.units.it/flora/?procedure=img_ins2&species_id=2881
Belgium, Han sur Lesse
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe
Growth form: Foliose, narrow lobed
Substrata: bark and rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: very common
Padanian area: rather rare
Humid submediterranean belt: very common
Humid mediterranean belt: rather common
Dry mediterranean belt: extremely rare
Predictive model
| Herbarium samples |
Index Fungorum
GBIF
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20462)
2001/12/05
detail of soredia
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20462)
2001/12/05
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10537)
2002/11/04
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli-Venezia Giulia, TS, Basovizza; altitude: 2
2008.02.21
with (in upper part of picture) Physconia distorta
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Slovenia, Lipica/Lipizza; altitude: 2
2008.02.21
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Slovenia, Lipica/Lipizza; altitude: 2
2008.02.21
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Slovenia, Lipica/Lipizza; altitude: 2
2008.02.21
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Slovenia, Lipica/Lipizza; altitude: 2
2008.02.21
Antonio Valero; Owner: Antonio Valero
Spain
2010
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste, Opicina, Viale alberato sulla strada per Monrupino
17/02/2017
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Felix Schumm – CC BY-SA 4.0
[10668], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, im Stubental ca. 5 km westlich von Sontheim, Straßenallee, ca.570 m, an Tilia. Leg et det. F. Schumm, 19.06.2003
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physconia_grisea.htm
France, 19/3/2013 - sur Acer - Mont Dore - Puy-de-Dôme
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physconia_grisea.htm
France, 19/3/2013 - sur Acer - Mont Dore - Puy-de-Dôme
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physconia_grisea.htm
France, 19/3/2013 - sur Acer - Mont Dore - Puy-de-Dôme
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Physconia_grisea.htm
France, 19/3/2013 - sur Acer - Mont Dore - Puy-de-Dôme
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1450&lang=en
France, Noirmoutier
Bernard Bouffinier - Source: https://italic.units.it/flora/?procedure=img_ins2&species_id=2881
France, Quimper
Bernard Bouffinier - Source: https://italic.units.it/flora/?procedure=img_ins2&species_id=2881
France, Quimper
Bernard Bouffinier - Source: https://italic.units.it/flora/?procedure=img_ins2&species_id=2881
France, Quimper
Jacques Haine - Source: https://italic.units.it/flora/?procedure=img_ins2&species_id=2881
Belgium, Han sur Lesse
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe
