Heterodermia subneglecta Elix

Australas. Lichenol., 68: 17, 2011.
Synonyms: Heterodermia japonica auct. eur.; Heterodermia obscurata auct. eur. p.max.p.; Polyblastidium subneglectum (Elix) Kalb
Distribution: N - Frl (Badin & Nimis 1996), Ven (Nascimbene & Caniglia 2000), TAA (B-Leg. H. Sipman nr. 8585), Emil (Fariselli & al. 2020), Lig (Petrak 1965, Castello & al. 1994, Brunialti & Giordani 2000, 2003, Giordani & al. 2001, 2002, Giordani 2006, Giordani & Incerti 2008, Watson 2014, Lich. Ital. Exs. 16: Isocrono & al. 2018). C - Tosc (Loppi & al. 1999a, Laganà & al. 2002, Lorenzini & al. 2003, Loppi & Frati 2006, Brunialti & Frati 2010, Brunialti & al. 2012b, Paoli & al. 2012, Benesperi & al. 2013), Umb (Ravera 2000, Panfili 2000, Ravera & al. 2006), Laz (Bartoli & al. 1997, Ravera 2001, Massari & Ravera 2002, Stofer 2006, Munzi & al. 2007), Mol (Frati & al. 2004, Caporale & al. 2008, Paoli & al. 2015), Sar (Zedda 2002). S - Camp (Nimis & Tretiach 2004, Nascimbene & al. 2010b, Brunialti & al. 2013, Ravera & Brunialti 2013), Cal (Puntillo 1996, Incerti & Nimis 2006).
Description: Thallus foliose, heteromerous, dorsiventral, narrow-lobed, loosely attached, forming irregular to orbicular, up to 5 cm wide rosettes. Lobes sublinear-elongate, radiating, widening towards the ascending lobe-tips, c. 2-3(-4) mm wide at tips, sometimes with lobules along the margins. Upper surface white to cream-coloured, glaucous when moist, sometimes pruinose at tips, with labriform soralia on lateral or terminal lobes. Lower surface white, arachnoid, spotted with an orange-red pigment, with marginal, simple or squarrosely branched, black, 1-3(-7) mm long rhizines. Upper cortex paraplectenchymatous; medulla white; lower cortex absent. Apothecia extremely rare, lecanorine, with a dark brown disc and a usually crenulate thalline margin. Epithecium brown; hymenium and hypothecium colourless. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, brown, broadly ellipsoid, 21-31 x 12-17 µm, Pachysporaria-type. Photobiont: chlorococcoid. Spot tests: cortex and medulla K+ yellow, C-, KC-, P- or P+ pale yellow; orange spots on lower surface K+ purple-violet. Chemistry: cortex with atranorin and chloroatranorin, medulla with zeorin, 6α-acetoxyhopane-16β,22-diol, and 7-chloroemodin.
Note: a mild-temperate species found on more or less isolated trees, occasionally on epilithic mosses; mainly Tyrrhenian, and also found in urban areas (e.g. at La Spezia). According to Roux & coll. (2014) most samples of Heterodermia obscurata from Europe do actually belong to this species, which, however, belong to a still poorly understood complex of mainly tropical species.
Growth form: Foliose, narrow lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: rather rare
Humid mediterranean belt: rather common
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm - CC BY-SA 4.0
[15873], Australia, Western Australia, Donnelly Well south of Yornup along SW Highway, wetland associated with Eucalyptus rudis and Melaleuca preissiana, 34.073° S, 116,180° E, 260 m. Leg. F. Schumm & E. Stocker-Wörgötter, 25.19 2009.



Felix Schumm - CC BY-SA 4.0
[15873], Australia, Western Australia, Donnelly Well south of Yornup along SW Highway, wetland associated with Eucalyptus rudis and Melaleuca preissiana, 34.073° S, 116,180° E, 260 m. Leg. F. Schumm & E. Stocker-Wörgötter, 25.19 2009.



Felix Schumm - CC BY-SA 4.0
[15873], Australia, Western Australia, Donnelly Well south of Yornup along SW Highway, wetland associated with Eucalyptus rudis and Melaleuca preissiana, 34.073° S, 116,180° E, 260 m. Leg. F. Schumm & E. Stocker-Wörgötter, 25.19 2009.



Felix Schumm - CC BY-SA 4.0
[15873], Australia, Western Australia, Donnelly Well south of Yornup along SW Highway, wetland associated with Eucalyptus rudis and Melaleuca preissiana, 34.073° S, 116,180° E, 260 m. Leg. F. Schumm & E. Stocker-Wörgötter, 25.19 2009.



Felix Schumm - CC BY-SA 4.0
[15873], Australia, Western Australia, Donnelly Well south of Yornup along SW Highway, wetland associated with Eucalyptus rudis and Melaleuca preissiana, 34.073° S, 116,180° E, 260 m. Leg. F. Schumm & E. Stocker-Wörgötter, 25.19 2009.


Felix Schumm - CC BY-SA 4.0
[15873], Australia, Western Australia, Donnelly Well south of Yornup along SW Highway, wetland associated with Eucalyptus rudis and Melaleuca preissiana, 34.073° S, 116,180° E, 260 m. Leg. F. Schumm & E. Stocker-Wörgötter, 25.19 2009
at: atranorin, z: zeorin, t2: japonin



Felix Schumm – CC BY-SA 4.0
[16263], Australia, New South Wales, Brown Mountain, near Cooma, Fred Piper Memorial Lookout, 36.597° S, 149.444° E, 910 m. Leg. F. Schumm & E. Stocker-Wörgötter, 13.11.2009.



Felix Schumm – CC BY-SA 4.0
[16263], Australia, New South Wales, Brown Mountain, near Cooma, Fred Piper Memorial Lookout, 36.597° S, 149.444° E, 910 m. Leg. F. Schumm & E. Stocker-Wörgötter, 13.11.2009.



Felix Schumm – CC BY-SA 4.0
[16263], Australia, New South Wales, Brown Mountain, near Cooma, Fred Piper Memorial Lookout, 36.597° S, 149.444° E, 910 m. Leg. F. Schumm & E. Stocker-Wörgötter, 13.11.2009.



Felix Schumm – CC BY-SA 4.0
[16263], Australia, New South Wales, Brown Mountain, near Cooma, Fred Piper Memorial Lookout, 36.597° S, 149.444° E, 910 m. Leg. F. Schumm & E. Stocker-Wörgötter, 13.11.2009.



Felix Schumm – CC BY-SA 4.0
[16263], Australia, New South Wales, Brown Mountain, near Cooma, Fred Piper Memorial Lookout, 36.597° S, 149.444° E, 910 m. Leg. F. Schumm & E. Stocker-Wörgötter, 13.11.2009.



Felix Schumm – CC BY-SA 4.0
[16263], Australia, New South Wales, Brown Mountain, near Cooma, Fred Piper Memorial Lookout, 36.597° S, 149.444° E, 910 m. Leg. F. Schumm & E. Stocker-Wörgötter, 13.11.2009.



Felix Schumm – CC BY-SA 4.0
[16263], Australia, New South Wales, Brown Mountain, near Cooma, Fred Piper Memorial Lookout, 36.597° S, 149.444° E, 910 m. Leg. F. Schumm & E. Stocker-Wörgötter, 13.11.2009.


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan


Bernard Bouffinier- Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=270&lang=en
France, Pointe de Dinan