Pseudevernia furfuracea (L.) Zopf var. furfuracea
Beih. Bot. Centralbl., 14: 124, 1903. Basionym: Lichen furfuraceus L. - Sp. Pl., 2: 1146, 1753.
Synonyms: Borrera furfuracea (L.) Ach.; Evernia furfuracea (L.) W. Mann; Parmelia furfuracea (L.) Ach.; Parmelia furfuracea f. elongata (Sambo) Zahlbr.; Parmelia furfuracea f. laricicola Gyeln.; Parmelia furfuracea var. cincinnata (Sambo) Zahlbr.; Parmelia soralifera (Bitter) Lynge; Pseudevernia soralifera (Bitter) Zopf
Distribution: N - VG (Castello 1996, 2002, Martellos 2003, Agnorelli & al. 2004, Martellos & Castello 2004, Castello & Skert 2005), Frl (Badin & Nimis 1996, Tretiach & Hafellner 2000, Martellos 2003, Castello & Skert 2005, Crisafulli & al. 2005, Tretiach & al. 2005, 2007, 2011, Rinino & al. 2005, Rinino 2006, Tretiach & Molaro 2007, Adamo & al. 2008, Bertuzzi & Tretiach 2013, Giordano & al. 2013), Ven (Caniglia & al. 1993b, 1999, Nimis 1994, Calliari & al. 1995, Nascimbene & Caniglia 1997, 2000b, 2002c, 2003c, Lazzarin 1997, 2000, Adamo & al. 2003b, Martellos 2003, Nascimbene 2005c, 2008, 2008c, 2011, Nascimbene & al. 2006e, 2007, 2009c, 2010b, 2014b, Nascimbene & Marini 2007, 2010, Brackel 2013, Kodnik & al. 2015), TAA (Follmann & Schulz 1993, Nascimbene & Caniglia 2000b, 2002c, Caniglia & al. 2002, Martellos 2003, Nascimbene 2001b, 2003, 2005b, 2006b, 2006c, 2008b, 2014, Gottardini & al. 2004, Nascimbene & al. 2005, 2006, 2006e, 2007b, 2009, 2010, 2014, 2014b, 2022, Stofer 2006, Brackel 2013, Lang 2009, Nascimbene & Marini 2015, Nimis & al. 2015, Trindade & al. 2021), Lomb (Rivellini 1994, Alessio & al. 1995, Arosio & Rinaldi 1995, Valcuvia & Gianatti 1995, Brusoni & al. 1997, Zocchi & al. 1997, Roella 1999, Brusoni & Valcuvia 2000, Casarini & al. 2000, Arosio & al. 2003, Martellos 2003, Valcuvia & al. 2003, Dalle Vedove & al. 2004, Nascimbene & al. 2006e, Bergamaschi & al. 2007, Furlanetto 2010, Brackel 2013, Gheza & al. 2015, 2020, 2023, Gheza 2019, 2019b), Piem (Caniglia & al. 1992, Morisi & Sereno 1995, Piervittori & al. 1997, Arosio & al. 1998, Piervittori 1998, 2003, Isocrono & Falletti 1999, Rizzio & al. 2001, Castino 2004, Isocrono & al. 2004, 2006, 2007, Martellos 2003, Isocrono & Ferrarese 2008, Isocrono & Piervittori 2008, Furlanetto 2010, Isocrono & al. 2011, Motiejunaitè & Grochowski 2014), VA (Borlandelli & al. 1996, Piervittori & Maffei 1996, 2001, Piervittori & Isocrono 1997, 1999, Bari & al. 2001, Piervittori & al. 2001, Revel & al. 2001, Ghiraldi 2003, Martellos 2003, Bergamaschi & al. 2004, Isocrono & al. 2005, Matteucci & al. 2008, 2008c, Isocrono & al. 2008, Loppi 2014, Malaspina & al. 2014, 2015, Casale & al. 2015, Lich. Ital. Exs. 42: Isocrono & al. 2020), Emil (Dalle Vedove & al. 2002, Martellos 2003, Benesperi 2009, Barba & al. 2012, Brackel 2015, Fariselli & al. 2020), Lig (Brunialti & al. 1999, Giordani & al. 2001, 2002, Brunialti & Giordani 2003, Martellos 2003, Giordani & Incerti 2008, Malaspina & al. 2009, 2014c). C - Tosc (Tretiach & Nimis 1994, Loppi & al. 1997, 1998, Loppi & Nascimbene 1998, Putortì & al. 1998, Tretiach & Ganis 1999, Paoli & Loppi 2001, Laganà & al. 2002, Martellos 2003, Benesperi & al. 2007, Benesperi & Lastrucci 2007, Lastrucci & al. 2009, Benesperi 2011, Brackel 2015, Nascimbene & al. 2015, Grifoni L. & al. 2021), Marc (Nimis & Tretiach 1999, Martellos 2003), Umb (Ravera 1998, Martellos 2003, Ravera & al. 2006, Panfili 2007, Brackel 2015), Laz (Owczarek & al. 2001, Ravera 2002, Martellos 2003, Guidotti & al. 2003, 2005, 2009, Massari & Ravera 2002, Ruisi & al. 2005, Guidotti & Owczarek 2013, Protano & al. 2014, 2015, Brackel 2015), Abr (Nimis & Tretiach 1999, Martellos 2003, Brackel 2015, Caporale & al. 2016, Gheza & al. 2021), Mol (Garofalo & al. 1999, Caporale & al. 2008, Brackel 2020), Sar (Zedda 1995, 2002, 2002b, Nöske 2000, Zedda & Sipman 2001, Martellos 2003, Cossu 2013). S - Camp (Garofalo & al. 1999, 2010, Aprile & al. 2002, 2003, 2003b, Martellos 2003, Basile & al. 2008, Nascimbene & al. 2010b, Brunialti & al. 2013, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Thüs & Licht 2006), Bas (Nimis & Tretiach 1999, Martellos 2003, Potenza 2006, Brackel 2011, Caggiano & al. 2015), Cal (Puntillo 1995, 1996, van den Boom & Giralt 2002, Martellos 2003, Incerti & Nimis 2006, Corapi & al. 2014, Gallo & al. 2014, Lucadamo & al. 2015, Brackel & Puntillo 2016), Si (Ottonello 1996, Czeczuga & al. 1999, Clocchiatti & al. 2000, 2002b, Martellos 2003, Merlo 2004, Grillo & Caniglia 2004, 2006, Ottonello 2005, Falco Scampatelli 2005, Iacolino & Ottonello 2006, Stofer 2006, Brackel 2008b, Carasci & Cataldo 2016, Campisi & al. 2020).
Description: Thallus foliose to subfruticose, of prostrate to ascending, flattened, strap-shaped 1-10(-20) mm wide, several cm long, dichotomously branched lobes. Upper surface grey to brown in exposed situations, matt, with cylindrical, simple to branched isidia, rarely also with orbicular soralia; lower surface usually channelled, black in older parts, white near the tips, without rhizines. Cortex paraplectenchymatous; medulla white, rather lax. Apothecia very rare, lecanorine, up to 15 mm across, more or less stalked, often irregular in form and sometimes with split margins, with a brown disc. Epithecium brown; hymenium and hypothecium colourless. Asci 8-spored, elongate-clavate, very thin-walled, with a K/I+ blue, tall tholus penetrated by a faintly amyloid apical cushion, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, ellipsoid, 7-10 x 4-6 µm. Pycnidia rare, immersed, black. Conidia sublageniform, 5-7 x c. 1 µm. Photobiont: chlorococcoid. Spot tests: cortex K+ yellow, C-, KC-, P-; medulla K-, C-, KC+ faintly pink, P-, UV-. Chemistry: cortex with atranorin and chloroatranorin, medulla with physodic acid.Note: a cool-temperate to boreal-montane lichen found on acid bark and lignum, occasionally also on siliceous rocks, with optimum in the montane and subalpine belts; abundant only in the Alps, rarer in the Apennines, exceptionally reaching the plains of Northern Italy on very acid substrata. References refer to the species in a broad sense; some records could refer to var. ceratea.
Growth form: Foliose, broad lobed
Substrata: rocks, bark, and lignum
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: extremely common
Oromediterranean belt: extremely rare
Montane belt: very common
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent
Predictive model
Herbarium samples
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (12146)
2001/11/29
var. soreumatica, detail of soredia
Andres Saag; Owner: University of Tartu
originally as Pseudevernia furfuracea
Triin Aimla; Owner: University of Tartu
originally as Pseudevernia furfuracea
P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36844)
2008.02.25
Tomi Trilar; Owner: Slovenian Museum of Natural History
Bulgaria, Blagoevgrad Province, Sandanski municipality, Slavyanka Mt. (Ali Botusch Mt.)
24.07,2008
Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)
with Pyrenula macrospora
Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)
Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)
Antonio Valero; Owner: Antonio Valero
Spain
2010
Giancarlo Medici; Owner: Giancarlo Medici
Italy, Modena, Appennino Modenese.
Herbarium: Forum Natura Mediterraneo
2010
Domenico Puntillo; Owner: Domenico Puntillo
Italy
Domenico Puntillo; Owner: Domenico Puntillo
Italy
Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Monte Bondone
2014
P.L. Nimis CC BY-SA 04
TSB 14009
J. Nascimbene CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
J. Nascimbene CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
J. Nascimbene CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
J. Nascimbene CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Hollergraben
2020
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Hollergraben
2020
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Hollergraben
2020
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Corse, L'Ospedale
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Corse, L'Ospedale
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Corse, Col de Bavella
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Corse, Col de Bavella
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Tartagine
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Tartagine
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Pays Baque, Col d'Iraty
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Mont Ventoux
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Mont Ventoux
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Mont Ventoux
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Mont Ventoux
Growth form: Foliose, broad lobed
Substrata: rocks, bark, and lignum
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: extremely common
Oromediterranean belt: extremely rare
Montane belt: very common
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent
Predictive model
| Herbarium samples |
Index Fungorum
GBIF
Juri Nascimbene; Owner: Department of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
03.04.2009
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (12146)
2001/11/29
var. soreumatica, detail of soredia
Andres Saag; Owner: University of Tartu
originally as Pseudevernia furfuracea
Triin Aimla; Owner: University of Tartu
originally as Pseudevernia furfuracea
P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (36844)
2008.02.25
Tomi Trilar; Owner: Slovenian Museum of Natural History
Bulgaria, Blagoevgrad Province, Sandanski municipality, Slavyanka Mt. (Ali Botusch Mt.)
24.07,2008
Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)
with Pyrenula macrospora
Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)
Gabriele Gheza, Tromello (PV); Owner: Gabriele Gheza, Tromello (PV)
Antonio Valero; Owner: Antonio Valero
Spain
2010
Giancarlo Medici; Owner: Giancarlo Medici
Italy, Modena, Appennino Modenese.
Herbarium: Forum Natura Mediterraneo
2010
Domenico Puntillo; Owner: Domenico Puntillo
Italy
Domenico Puntillo; Owner: Domenico Puntillo
Italy
Juri Nascimbene - CC BY-SA 4.0; Owner: ITALIC - Dyades Project - Dept. of Life Sciences, University of Trieste
Italy, Trentino-Alto Adige, Trento, Monte Bondone
2014
P.L. Nimis CC BY-SA 04
TSB 14009
J. Nascimbene CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
J. Nascimbene CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
J. Nascimbene CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
J. Nascimbene CC BY-SA 4.0
Italy, Trentino-Alto Adige, Trento, Villa Welsperg, Centro Visitatori del Parco di Paneveggio; altitude: 1030 m
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Hollergraben
2020
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Hollergraben
2020
Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Hollergraben
2020
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Corse, L'Ospedale
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Corse, L'Ospedale
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Corse, Col de Bavella
Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Corse, Col de Bavella
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Tartagine
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Tartagine
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Pays Baque, Col d'Iraty
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Mont Ventoux
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Mont Ventoux
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=611&lang=en
France, Mont Ventoux
