Pyrenodesmia chalybaea (Fr.) A. Massal.

Atti reale Ist. Veneto Sci. ser. 2., 3 append.: 123, 1853. Basionym: Parmelia chalybaea Fr. - Lichenogr. Eur. Ref.: 123, 1831.
Synonyms: Callopisma variabile var. lilacinum (A. Massal.) Müll. Arg.; Caloplaca chalybaea (Fr.) Müll. Arg.; Caloplaca olivacea (A. Massal.) Jatta; Caloplaca variabilis f. chalybaea (Fr.) Clauzade & Cl. Roux; Caloplaca variabilis var. lilacina (A. Massal.) Jatta; Caloplaca variabilis var. ocellulata (Ach.) Boistel; Caloplaca variabilis var. ocellulata f. chalybaea (Fr.) Clauzade & Cl. Roux; Catillaria lapsans (Nyl.) Boistel; Placodium chalybaeum (Fr.) Hepp; Pyrenodesmia olivacea A. Massal.
Distribution: N - VG, Frl, Ven (Lazzarin 2000b, Nascimbene & Caniglia 2003c, Nascimbene & Marini 2007, Watson 2014), TAA (Nascimbene & al. 2022), Lomb (Florio & al. 2004, 2006), Piem (TSB 32936), VA (Piervittori & Isocrono 1999), Emil (Fariselli & al. 2020), Lig (Giordani & al. 2016). C - Tosc, Marc (Nimis & Tretiach 1999), Umb (Nimis & Tretiach 1999, Ravera & al. 2006), Laz (Nimis & Tretiach 2004), Abr (Nimis & Tretiach 1999, Gheza & al. 2021), Mol (Nimis & Tretiach, 1999, 2004, Caporale & al. 2008, Genovesi & Ravera 2014), Sar (Brackel & Berger 2019). S - Camp (Garofalo & al. 1999, 2010, Aprile & al. 2003b, Nimis & Tretiach 2004), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999), Cal (Puntillo 2011), Si (Nimis & al. 1994, 1996b, Schicchi & al. 1997, Grillo 1998, Grillo & Caniglia 2004, Campisi & al. 2020).
Description: Thallus crustose, episubstratic, areolate, usually pale grey with a bluish tinge, more rarely dark grey to grey-brown, often slightly pruinose, sometimes delimited by a dark prothallus, forming neat orbicular patches to a few cm across. Areoles distinctly flat, the peripheral ones often slightly elongate and separated by radiating thin cracks, giving the thallus a subplacodioid appearance. Apothecia lecanorine, 0.3-0.7 mm across, immersed in the areoles, with a usually flat, sometimes slightly pruinose, black disc, and a white or blue-white pruinose, thin, persistent thalline margin. Proper exciple poorly developed, brown in outer part, colourless to brownish within, the pigmented parts K+ violet; epithecium grey to brown, K+ violet; hymenium colourless, without crystals; paraphyses c. 1.5 μm thick at base, the apical cells up to 4-5 μm wide; hypothecium colourless. Asci 8-spored, clavate, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 2-celled, polarilocular, hyaline, ellipsoid, (9-)12-15 x 5-8 μm, the equatorial thickening (“septum”) 3-5 μm, c. 1/3 of spore length. Photobiont chlorococcoid. Spot tests: thallus K- or K+ faintly violet, C-, KC-, P-; pigmented parts of apothecia K+ violet and sometimes N+ pale violet. Chemistry: Sedifolia-grey pigment in apothecia and sometimes in thallus.
Note: a mild-temperate species known from Europe and adjoining Africa and Asia, found on hard calciferous rocks (mostly on compact limestone) and dolomite, often, but not exclusively, on steeply inclined faces; especially common in Central and Southern Italy, above the Mediterranean belt.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rare
Subalpine belt: rather rare
Oromediterranean belt: extremely common
Montane belt: extremely common
Submediterranean belt: rather common
Padanian area: absent
Humid submediterranean belt: rather common
Humid mediterranean belt: rather rare
Dry mediterranean belt: very rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum Bílé Karpaty dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.


Department of Life Sciences, University of Trieste
2001/11/12


Manilio Prignano; Owner: Manilio Prignano
Italia, M. Navegna, Appennino Centrale
2010


Manilio Prignano; Owner: Manilio Prignano
Italia, M. Navegna, Appennino Centrale
2010


Manilio Prignano; Owner: Manilio Prignano
Italia, M. Navegna, Appennino Centrale
2010



Domenico Puntillo; Owner: Domenico Puntillo
Italy


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php (Courtesy: Anita Stridvall)



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017



Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum Bílé Karpaty dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum Bílé Karpaty dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum Bílé Karpaty dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19572], Bohemoslovacia, Slovacia, jugum Carpatorum 'Bílé Karpaty' dictum, in monte Vršatec, alt. 850 m, ad saxa calcarea. Leg. I. Pišút et A. Vezda, 31.05.1971. VEZDA: LICHENES SELECTI EXSICCATI NR. 1145.



Felix Schumm – CC BY-SA 4.0
[17256],Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, 0.7 km östlich von Dunstelkingen, sonnige Kalkfelsen, 48.718657° N, 10.426030° E, 530 m. Leg. et det. Schumm 13.05.2011.


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Jennifer Fiorentino
Malta


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Pyrenodesmia_chalybaea.htm
France, session AFL Luberon 2009 - Vaucluse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Pyrenodesmia_chalybaea.htm
France, session AFL Luberon 2009 - Vaucluse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Pyrenodesmia_chalybaea.htm
France, session AFL Luberon 2009 - Vaucluse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_5/Pyrenodesmia_chalybaea.htm
France, session AFL Luberon 2009 - Vaucluse


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Salcedo, ermita del Alba (Quirós-Asturias), 29-XI-22, sobre rocas calizas, junto a Variospora placidia.