Pyrenula nitidella (Schaer.) Müll. Arg.

in Engler, Bot. Jahrb., 6: 414, 1885. Basionym: Verrucaria nitida var. nitidella Flörke ex Schaer. - Lich. Helv. Spicil., 2: 58, 1826.
Synonyms: Bunodea nitida var. nitidella (Schaer.) Beltr.; Pyrenula nitida var. dermatodes (Borrer) Trevis.; Pyrenula nitida var. nitidella (Schaer.) Schaer.
Distribution: N - Ven (Ravera & al. 2021b), Lomb (Ravera & al. 2021b), VA (Piervittori & Isocrono 1999), Emil (Fariselli & al. 2020), Lig (Giordani & al. 2002, Brunialti & Giordani 2003, Giordani & Incerti 2008, Watson 2014). C - Tosc (Senese & Critelli 2000, Loppi & al, 2004c), Laz (Ravera 2006c, Munzi & al. 2012), Sar (Zedda & al. 2001, Zedda 2002). S - Camp, Pugl (Thüs & Licht 2006), Cal (Puntillo 1996).
Description: Thallus crustose, more or less immersed in the bark, continuous, smooth, olive-brown to pale brown, often with scattered, whitish, punctiform pseudocyphellae. Perithecia black, 0.2-0.35 mm across, numerous, hemispherical to broadly conical, forming low projections. Exciple dark brown throughout, containing colourless crystals; hymenium colourless, not inspersed with oil droplets, the part adjacent to exciple with brown-orange crystals of anthraquinones reacting K+ red; hamathecium of branched and anastomosed periphysoids, later substituted by more or less unbranched paraphyses; ostiole with periphyses. Asci 8-spored, narrowly cylindrical, long-stalked, bitunicate, with tholus, thickened at apex with an internal apical beak, non-amyloid. Ascospores 3-septate, brown, thick-walled, narrowly ellipsoid, with 4 diamond-shaped locules, (20-)22-26(-28) x (8-)8.5-11(-12) μm. Photobiont trentepohlioid. Spot tests: thallus: K+ yellow to red-orange around perithecia, C-, KC-, P-, UV-. Chemistry: anthraquinones in perithecia and sometimes in thallus.
Note: a mild-temperate to Mediterranean-Atlantic species found on the bark of deciduous trees in open, humid woodlands; very rare in the North, more frequent in Tyrrhenian Italy.
Growth form: Crustose
Substrata: bark
Photobiont: Trentepohlia
Reproductive strategy: mainly sexual
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: rare
Humid mediterranean belt: very rare
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm - CC BY-SA 4.0
[4223], Germany, Baden-Württemberg, Kreis Schwäbisch Gmünd, im Beutenbachtal bei der Beutenmühle ca. 3 km südöstlich von Lorch, ca 310 m, an Fraxinus, TK 7224. Leg. et det. Schumm 29.05.1995



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (4184)
2001/11/29


Andres Saag; Owner: University of Tartu



Felix Schumm - CC BY-SA 4.0
[4223], Germany, Baden-Württemberg, Kreis Schwäbisch Gmünd, im Beutenbachtal bei der Beutenmühle ca. 3 km südöstlich von Lorch, ca 310 m, an Fraxinus, TK 7224. Leg. et det. Schumm 29.05.1995



Felix Schumm - CC BY-SA 4.0
[4223], Germany, Baden-Württemberg, Kreis Schwäbisch Gmünd, im Beutenbachtal bei der Beutenmühle ca. 3 km südöstlich von Lorch, ca 310 m, an Fraxinus, TK 7224. Leg. et det. Schumm 29.05.1995



Felix Schumm - CC BY-SA 4.0
[4223], Germany, Baden-Württemberg, Kreis Schwäbisch Gmünd, im Beutenbachtal bei der Beutenmühle ca. 3 km südöstlich von Lorch, ca 310 m, an Fraxinus, TK 7224. Leg. et det. Schumm 29.05.1995



Felix Schumm - CC BY-SA 4.0
[4223], Germany, Baden-Württemberg, Kreis Schwäbisch Gmünd, im Beutenbachtal bei der Beutenmühle ca. 3 km südöstlich von Lorch, ca 310 m, an Fraxinus, TK 7224. Leg. et det. Schumm 29.05.1995



Felix Schumm - CC BY-SA 4.0
[4223], Germany, Baden-Württemberg, Kreis Schwäbisch Gmünd, im Beutenbachtal bei der Beutenmühle ca. 3 km südöstlich von Lorch, ca 310 m, an Fraxinus, TK 7224. Leg. et det. Schumm 29.05.1995



Felix Schumm - CC BY-SA 4.0
[4223], Germany, Baden-Württemberg, Kreis Schwäbisch Gmünd, im Beutenbachtal bei der Beutenmühle ca. 3 km südöstlich von Lorch, ca 310 m, an Fraxinus, TK 7224. Leg. et det. Schumm 29.05.1995



Felix Schumm - CC BY-SA 4.0
[4223], Germany, Baden-Württemberg, Kreis Schwäbisch Gmünd, im Beutenbachtal bei der Beutenmühle ca. 3 km südöstlich von Lorch, ca 310 m, an Fraxinus, TK 7224. Leg. et det. Schumm 29.05.1995



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50



Felix Schumm CC BY-SA 4.0
13\50


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1055&lang=en
France, Crozon
with P. chlorospila


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1055&lang=en
France, Crozon
with P. chlorospila, which reacts UV+ white


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1055&lang=en
France, Crozon


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1055&lang=en
France, Crozon


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1055&lang=en
France, Crozon


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1055&lang=en
France, Crozon


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse