Rinodina immersa (Körb.) J. Steiner

Sitzungsber. Akad. Wiss. Wien, math.-naturw. Kl., Abt. 1, 102: 164, 1893. Basionym: Rinodina bischoffii var. immersa Körb. - Parerga Lichenol.: 75, 1859.
Synonyms: Rinodina bischoffii var. exigua Müll. Arg.; Rinodina bischoffii var. intermedia Müll. Arg.; Rinodina bischoffii var. ochracea Müll. Arg.; Rinodina bischoffii var. perexigua Müll. Arg.
Distribution: N - VG (Nimis & Tretiach 1995, Tretiach & Pecchiari 1995, Geletti 1997, Pinna & al. 1998, Crisafulli & al. 2006, Piervittori & al. 2006, Tretiach & al. 2007b), Frl (TSB 3005), Ven (Nascimbene & Caniglia 2000, 2003c), TAA (Spitale & Nascimbene 2012, Nascimbene & al. 2022), Lomb (Ravera & al. 2023b), Lig (TSB 33575). C - Tosc, Marc (Nimis & Tretiach 1999), Umb (Nimis & Tretiach 1999, Ravera & al. 2006), Laz (Nimis & Tretiach 2004), Abr (Nimis & Tretiach 1999, Cucchi & al. 2009, Caporale & al. 2016, Gheza & al. 2021), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Ravera & al. 2009, Brackel 2020), Sar (Rizzi & al. 2011). S - Camp (Aprile & al. 2003b, Nimis & Tretiach 2004, Garofalo & al. 2010), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si (Nimis & al. 1996b, Grillo 1998, Grillo & Caniglia 2004, Brackel 2008b, 2008c).
Description: Thallus crustose, endosubstratic, or very thinly episubstratic and then whitish to grey, usually poorly evident. Apothecia 0.2-0.6(-0.8) mm across, cryptolecanorine, immersed in pits in the rock, with a flat, usually epruinose, brown to black disc and a thin, concolorous or paler proper margin; thalline margin absent. Proper exciple brown in outer part, colourless within; epithecium dark brown, up to 20 µm high, K-; hymenium colourless, 80-100 µm high, not inspersed with oil droplets; paraphyses mostly simple, 1.5-2.5 μm thick at mid-level, the apical cells 5-8 μm wide, with an internal brown pigment cap; hypothecium colourless. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, brown, ellipsoid, (13-)16-22 x 9-13 μm, Bischoffii-type, with a broad pigmented band around septum, the torus not developed, the ontogeny of type A (apical wall thickening after septum formation). Pycnidia rare. Conidia colourless, bacilliform, c. 4 x 1 μm. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a mainly southern species found on horizontal to steeply inclined, dry surfaces of limestone and (more rarely) dolomite wetted by rain, but also on pebbles, exceptionally reaching the Alpine belt.
Growth form: Crustose endolithic
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: very rare
Subalpine belt: rare
Oromediterranean belt: rare
Montane belt: rather common
Submediterranean belt: very common
Padanian area: very rare
Humid submediterranean belt: common
Humid mediterranean belt: rather common
Dry mediterranean belt: rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis CC BY-SA 04
TSB 38056



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11341)
2001/11/29



Domenico Puntillo; Owner: Department of Life Sciences, University of Trieste
Italy, Calabria



Domenico Puntillo; Owner: Department of Life Sciences, University of Trieste
Italy, Calabria



Felix Schumm - CC BY-SA 4.0
[19553], Italia, Abruzzo, prov. L'Aquila, loco Rovere, prope ruinam arcis, 1440 m, ad saxa calcarea. Leg. M. Tretiach & D. Pinna (TSB 34131, holotyous), 23.6.2003. VEZDA: LICHENES RARIORES EXSICCATI



Domenico Puntillo; Owner: Department of Life Sciences, University of Trieste
Italy, Calabria



Felix Schumm - CC BY-SA 4.0
[19553], Italia, Abruzzo, prov. L'Aquila, loco Rovere, prope ruinam arcis, 1440 m, ad saxa calcarea. Leg. M. Tretiach & D. Pinna (TSB 34131, holotyous), 23.6.2003. VEZDA: LICHENES RARIORES EXSICCATI



Felix Schumm - CC BY-SA 4.0
[19553], Italia, Abruzzo, prov. L'Aquila, loco Rovere, prope ruinam arcis, 1440 m, ad saxa calcarea. Leg. M. Tretiach & D. Pinna (TSB 34131, holotyous), 23.6.2003. VEZDA: LICHENES RARIORES EXSICCATI



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001



Felix Schumm; Owner: CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens



Felix Schumm - CC BY-SA 4.0
[TSB_34131], Italia, Abruzzo, Prov. L’Aquila, loc. Rovere, 1440 m, on limestone. Leg. M. Tretiach & D. Pinna, 23.06.2001. with Pyrenodesmia erodens


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm



Pier Luigi Nimis - CC BY-SA 4.0
TSB 37854



Pier Luigi Nimis - CC BY-SA 4.0
TSB 37854



Felix Schumm - CC BY-SA 4.0
[19553], Italia, Abruzzo, prov. L'Aquila, loco Rovere, prope ruinam arcis, 1440 m, ad saxa calcarea. Leg. M. Tretiach & D. Pinna (TSB 34131, holotyous), 23.6.2003. VEZDA: LICHENES RARIORES EXSICCATI



P.L. Nimis CC BY-SA 4.0
TSB 38056



P.L. Nimis CC BY-SA 4.0
TSB 38056



Felix Schumm - CC BY-SA 4.0
[19553], Italia, Abruzzo, prov. L'Aquila, loco Rovere, prope ruinam arcis, 1440 m, ad saxa calcarea.
Leg. M. Tretiach & D. Pinna (TSB 34131, holotyous), 23.6.2003. VEZDA: LICHENES RARIORES EXSICCATI



P.L. Nimis CC BY-SA 4.0
TSB 38056



P.L. Nimis CC BY-SA 04
TSB 38056



P.L. Nimis CC BY-SA 04
TSB 38056


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R/Rinodina_immersa.htm
France, 10/7/2015 - session AFL Bourgogne - Côte-d'Or


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R/Rinodina_immersa.htm
France, 10/7/2015 - session AFL Bourgogne - Côte-d'Or


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R/Rinodina_immersa.htm
France, 10/7/2015 - session AFL Bourgogne - Côte-d'Or


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R/Rinodina_immersa.htm
France, 10/7/2015 - session AFL Bourgogne - Côte-d'Or