Rinodina milvina (Wahlenb.) Th. Fr.

N. Acta Reg. Soc. Sci. Upsal., ser. 3, 3: 224, 1861. Basionym: Parmelia milvina Wahlenb. in Ach. - Meth. Lich. Suppl.: 34, 1803.
Synonyms: Lecanora milvina (Wahlenb.) Ach.; Lecanora sophodes var. scopulina Nyl.; Lecanora sophodes var. submilvina Nyl.; Lecanora subconfragosa Nyl.; Rinodina milvina var. karelica Räsänen; Rinodina milvina var. scopulina (Nyl.) H. Olivier; Rinodina sophodes f. saxicola Kernst.; Rinodina sophodes var. scopulina (Nyl.) Croz.; Rinodina subconfragosa (Nyl.) Flagey
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven, TAA (Nascimbene 2003, Nascimbene & al. 2022), Lomb, Piem (Isocrono & al. 2004, Favero-Longo & al. 2015), VA (Piervittori & Isocrono 1999, Matteucci & al. 2008c, 2015c, Favero-Longo & Piervittori 2009, Isocrono & al. 2008), Emil (Tretiach & al. 2008, Fariselli & al. 2020), Lig (Brunialti & al. 1999). C - Tosc, Sar (Nöske 2000, Rizzi & al. 2011, Giordani & al. 2013). S - Pugl, Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996), Si.
Description: Thallus crustose, episubstratic, rimose-areolate, often delimited by a black prothallus, the areoles up to 0.5-1 mm wide, smooth to rugose, dark grey, reddish brown or dark brown, dull. Medulla white, I-. Apothecia lecanorine, adnate to subsessile, contiguous, sometimes angular by mutual compression, 0.3-1(-1.5) mm across, with a dark brown to black, concave to flat, rarely slightly convex disc and a thick, smooth to crenulate, persistent thalline margin. Thalline exciple 60-100 µm wide laterally, corticate, without crystals; proper exciple colourless, 5-20 µm wide laterally, expanding to 10-40 µm in upper part; epithecium reddish brown, K-; hymenium colourless, (65-)100-120 µm high; paraphyses not strongly coherent, 1.5-2.5 µm thick at mid-level, the apical cells 3.5-5.5 µm wide, immersed in dispersed pigment; hypothecium colourless. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, often constricted at septum, brown, broadly ellipsoid, (13-)16-19(-23) x (7-)9-11(-13) µm, Milvina-type, often grading into Pachysporaria-type at maturity, with a well-developed torus, the walls finely warted, the ontogeny of type A (apical wall thickening after septum formation). Pycnidia dark, immersed. Conidia bacilliform, 4-5 x 1-1.5 µm. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a cool-temperate to arctic-alpine, circumpolar lichen found on boulders of siliceous rocks, usually on steeply inclined surfaces, often (but not always) parasitic on other crustose lichens; the Italian distribution ranges from the Alps to the high Mediterranean mountains, with optimum near or above treeline.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
paras crustose lichens when young

Commonnes-rarity: (info)

Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: rather common
Montane belt: rather rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples
INDEX FUNGORUM GBIF DOLICHENS


Pier Luigi Nimis - CC BY-SA 4.0
TSB 42444



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34912)
2002/02/21



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20168)
2001/11/29


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm


Harrie Sipman - Source: https://archive.bgbm.org/sipman/Zschackia/AegeanLichens/Rinodina.htm



P.L. Nimis CC BY-SA 4.0
TSB 42444



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34912)
2002/02/21


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R2/Rinodina_milvina.htm
France, 2/9/2014 - Bonneval-sur-Arc, Pont de L'Oulietta, alt. 2500 m - Savoie


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_R/Textes_R2/Rinodina_milvina.htm
France, 2/9/2014 - Bonneval-sur-Arc, Pont de L'Oulietta, alt. 2500 m - Savoie



Felix Schumm - CC BY-SA 4.0
[VZR156}, Austria. Carinthia, pars Alpinum Gurktaler Alpen dictum, ad latera austro-orientalia montis Gregerlnock, 1850-2290 m. Ad saxa schistosa. Leg. R. Türk & H. Wittman, 21.9.1988. EX A. VEZDA: LICHENES RARIORES EXSICCATI NR. 156.



Felix Schumm - CC BY-SA 4.0
[VZR156}, Austria. Carinthia, pars Alpinum Gurktaler Alpen dictum, ad latera austro-orientalia montis Gregerlnock, 1850-2290 m. Ad saxa schistosa. Leg. R. Türk & H. Wittman, 21.9.1988. EX A. VEZDA: LICHENES RARIORES EXSICCATI NR. 156.



Felix Schumm - CC BY 4.0
[VZ1748], Italia.Basilicata, Prov. Potenza: Monte Pollino, Vacquarro, 1460 m. Ad saxa siliceo-calcarea. Leg. H. Mayrhofer, 2.7.1980. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 1748.



Felix Schumm - CC BY 4.0
[VZ1748], Italia.Basilicata, Prov. Potenza: Monte Pollino, Vacquarro, 1460 m. Ad saxa siliceo-calcarea. Leg. H. Mayrhofer, 2.7.1980. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 1748.



Felix Schumm - CC BY 4.0
[VZ1861], URSS. Armenia, distr. Krasnosirijsk: ad latera merid. montium Sevanskij xhrebret, supra lacum Sevan dictutm, 000 m. Ad saxa vulcanica. Leg. A. Vezda et V. Vašák, 6.7.1983, det H. Mayrhofer. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 1861



Felix Schumm - CC BY 4.0
[VZ1861], URSS. Armenia, distr. Krasnosirijsk: ad latera merid. montium Sevanskij xhrebret, supra lacum Sevan dictutm, 000 m. Ad saxa vulcanica. Leg. A. Vezda et V. Vašák, 6.7.1983, det H. Mayrhofer. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 1861



Felix Schumm - CC BY 4.0
[VZ2297], Austria. Tirolia, alpes dicti "Hohe Tauern": Gruben, secus viam inter Stein et Casas alpinas Sudetendeutsche Hütte dictas, 1800 m. Ad saxa Schistosa. Leg. J. Horáková et A. Vezda. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 2297



Felix Schumm - CC BY 4.0
[VZ2297], Austria. Tirolia, alpes dicti "Hohe Tauern": Gruben, secus viam inter Stein et Casas alpinas Sudetendeutsche Hütte dictas, 1800 m. Ad saxa Schistosa. Leg. J. Horáková et A. Vezda. EX A. VEZDA LICHENES SELECTI EXSICCATI NR. 2297


Source: Giralt M. & Llimona X. (1997) Mycotaxon 52: 175-224