Rinodinella controversa (A. Massal.) H. Mayrhofer & Poelt

Rinodinella controversa (A. Massal.) H. Mayrhofer & Poelt

Hoppea, 37: 92, 1978. Basionym: Rinodina controversa A. Massal. - Ric. Auton. Lich. Crost.: 16, 1852.

Synonyms: Berengeria fusca (A. Massal.) Trevis.; Buellia dubyana var. nigrescens Müll. Arg.; Catolechia fusca A. Massal.; Lecanora budensis Nyl.; Lecanora crustulata (A. Massal.) Stizenb.; Lecanora sophodes. f. controversa (A. Massal.) Nyl.; Rinodina budensis (Nyl.) Zahlbr.; Rinodina controversa f. crustulata A. Massal.; Rinodina crustulata (A. Massal.) Arnold; Rinodina fusca (A. Massal.) Bagl.; Rinodina sublobata (Arnold) H. Olivier

Distribution: N - VG, Frl, Ven (Lazzarin 2000b), TAA (Nascimbene & al. 2022), Lomb, Piem (Isocrono & al. 2004), VA (Piervittori & Isocrono 1999), Emil (Fariselli & al. 2020), Lig. C - Tosc (Benesperi 2000a, 2006), Umb (Genovesi & Ravera 2001, Ravera & al. 2006), Laz (Roccardi 2003), Abr (Nimis & Tretiach 1999), Mol (Nimis & Tretiach 1999, Caporale & al. 2008), Sar. S - Camp (Garofalo & al. 1999, 2010, Aprile & al. 2003b), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999), Cal (Puntillo 2011), Si (Nimis & al. 1996b, Grillo 1998, Grillo & Caniglia 2004, Grillo & al. 2007b).

Description: Thallus crustose, episubstratic, areolate, brown to dark brown, almost black in sun-forms, epruinose, rarely delimited by a brown prothallus, often covering relatively large surfaces, the areoles contiguous, subangular, flat, 0.2-0.8 mm wide. Cortex brown in upper part, colourless in lower part, formed of anticlinally arranged hyphae; medulla white, I-. Apothecia lecanorine or cryptolecanorine, sessile, (0.5-)0.8-1(-1.3) mm across, with a flat to convex, dark brown to black, epruinose disc, a thin, smooth, finally rarely excluded thalline margin, and sometimes a thin parathecial ring within the thalline margin. Proper exciple of radiating hyphae; epithecium reddish brown, K-; hymenium colourless, sometimes with oil droplets, up to100 μm high, K/I+ blue; paraphyses branched in upper part, c. 1.2 μm thick at mid-level, the apical cells 3-5 μm wide, with a thin internal cap of brown pigment; hypothecium colourless. Asci 8-spored, clavate, the K/I+ blue tholus penetrated by a faintly amyloid apical cushion with parallel or diverging flanks, the wall K/I-, surrounded by a K/I+ blue outer layer, Lecanora-type. Ascospores 1-septate, pale brown, ellipsoid but often slightly constricted at septum, thin-walled throughout, 13-20 x 5-7 μm, Rinodinella-type, the septum very thin (1 μm or less), with ontogeny of type A. Pycnidia immersed, the wall colourless. Conidia bacilliform to narrowly ellipsoid, 4-5 x c. 1 µm. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.

Note: a mainly southern species in Europe, found on the top of exposed calcareous boulders, with optimum below the montane belt.

Growth form: Crustose

Substrata: rocks

Photobiont: green algae other than Trentepohlia

Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent

Subalpine belt: absent

Oromediterranean belt: absent

Montane belt: rare

Submediterranean belt: rather common

Padanian area: extremely rare

Humid submediterranean belt: rather common

Humid mediterranean belt: rather rare

Dry mediterranean belt: very rare

pH of the substrata:

1

2

3

4

5

(info)

Solar irradiation:

1

2

3

4

5

(info)

Aridity:

1

2

3

4

5

(info)

Eutrophication:

1

2

3

4

5

(info)

Poleotolerance:

0

1

2

3

(info)

Altitudinal distribution:

1

2

3

4

5

6

(info)

Predictive model

Herbarium samples

P.L. Nimis CC BY-SA 4.0

TSB 2307

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste

Herbarium: TSB (2307)
2001/12/10

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste

Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste

Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste

Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste

Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste

Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017

Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste

Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017

Andrea Moro; Owner: Department of Life Sciences, University of Trieste

italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017

Andrea Moro; Owner: Department of Life Sciences, University of Trieste

italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017

Andrea Moro; Owner: Department of Life Sciences, University of Trieste

italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017

Andrea Moro; Owner: Department of Life Sciences, University of Trieste

italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017

Andrea Moro; Owner: Department of Life Sciences, University of Trieste

italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017

Andrea Moro; Owner: Department of Life Sciences, University of Trieste

italy, Friuli Venezia Giulia, Trieste, Trieste Karst near Borgo Grotta Gigante
16/02/2017

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste

Herbarium: TSB (2307)
2001/12/10

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