Scytinium lichenoides (L.) Otálora, P.M. Jørg. & Wedin
Fungal Divers., 64, 1: 290, 2013. Basionym: Tremella lichenoides L. - Sp. Pl.: 1157, 1753.
Synonyms: Collema atrocoeruleum (Schaer.) Rabenh.; Collema fimbriatum (Ach.) Röhl.; Collema lacerum DC.; Leptogium atrocoeruleum (Schaer.) A. Massal.; Leptogium lacerum (Sw.) Gray; Leptogium lacerum var. majus Körb.; Leptogium lacerum var. sendtneri (Flot.) Müll. Arg.; Leptogium lichenoides (L.) Zahlbr.; Leptogium lophaeum (Ach.) Cromb.; Leptogium pellucidum M. Choisy
Distribution: N - VG, Frl (Tretiach & Hafellner 2000, Tretiach & Molaro 2007, Tomasi 2007, Otálora & al. 2008, Brackel 2013), Ven (Caniglia & al. 1999, Nascimbene & Caniglia 2003c, Nascimbene 2005c, 2008c, 2011, Nascimbene & al. 2005b, 2006c, 2007, 2010b, Thor & Nascimbene 2007, Nascimbene & Marini 2007, Otálora & al. 2008), TAA (Philippi 1983, De Benetti & Caniglia 1993, Nascimbene 2003, 2005b, 2008b, Nascimbene & al. 2006, 2007b, 2022, Nimis & al. 2015), Lomb (Philippi 1983, Gheza 2019, 2019b, Gheza & al. 2023), Piem (Isocrono & al. 2004), VA (Piervittori & Isocrono 1999), Emil (Nimis & al. 1996, Fariselli & al. 2020), Lig (Brunialti & al. 1999, Valcuvia & al. 2000, Otálora & al. 2004, Giordani & al. 2016, Ravera & al. 2022). C - Tosc (Tretiach & Nimis 1994, Otálora & al. 2004, Benesperi 2006, 2011, Benesperi & al. 2007, Brackel 2015), Marc (Nimis & Tretiach 1999), Umb (Ravera 1998, Nimis & Tretiach 1999, Ravera & al. 2006, Panfili 2007), Laz (Ravera 2002, Massari & Ravera 2002, Nimis & Tretiach 2004, Ravera & Genovesi 2008 Otálora & al. 2008, Brackel 2015), Abr (Nimis & Tretiach 1999, Brackel 2015, Corona & al. 2016, Di Nuzzo & al. 2021, Gheza & al. 2021, Vallese & al. 2022), Mol (Garofalo & al. 1999, Nimis & Tretiach, 1999, 2004, Caporale & al. 2008, Genovesi & Ravera 2014, Brackel 2020), Sar (Zedda 2002, 2002b, Zedda & al. 2010, Rizzi & al. 2011, Cogoni & al. 2011, Di Nuzzo & al. 2022). S - Camp (Garofalo & al. 1999, 2010, Aprile & al. 2002, 2003, 2003b, Nimis & Tretiach 2004, Nascimbene & al. 2010b, Brunialti & al. 2013, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Garofalo & al. 1999, Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999, Potenza 2006, Brackel 2011), Cal (Puntillo 1996, Puntillo & Puntillo 2004, Incerti & Nimis 2006, Stofer 2006), Si (Nimis & al. 1994, 1996b, Ottonello & Salone 1994, Ottonello & al. 1994, 2011, Ottonello 1996, Ottonello & Romano 1997, Grillo 1998, Grillo & Caniglia 2004, Merlo 2004, 2004b, Brackel 2008b Otálora & al. 2008, Gianguzzi & al. 2009, Liistro & Cataldo 2011, Campisi & al. 2020).
Description: Thallus minutely foliose to squamulose, slightly gelatinous when wet, of ascending, crowded lobes, often forming 2-4(-6) cm wide, loose cushions. Lobes grey-brown to dark brown, often paler and bluish grey in the basal, less exposed parts, thin, loosely attached, (0.3-)1-3(-4) mm broad, up to c. 1 cm long, 80-150 µm thick, more or less imbricate, the surface wrinkled, the edges deeply dilacerate-fimbriate, with cylindrical, isidia-like, often dense marginal outgrowths. Lower surface paler, erhizinate, with distinct, raised, longitudinal ribs and scattered white hapters. Upper and lower cortex of a single layer of angular, more or less isodiametrical cells, the layer inbetween of intertwined hyphae and loosely arranged chains of Nostoc. Apothecia rare, lecanorine, strongly constricted at base, 0.3-0.7(-1) mm across, with a brown, slightly concave to flat disc, and a thalline margin often covered in coralloid outgrowths. Proper exciple euparaplectenchymatous, 20-70 µm wide laterally; epithecium brownish; hymenium colourless, 130-200 µm high, I+ blue; paraphyses coherent, mostly simple, 1-1.5 μm thick at mid-level, the apical cells slightly swollen; hypothecium pale yellow, 35-70 µm high. Asci 8-spored, cylindrical-clavate, the apex strongly thickened, the apical dome K/I+ pale blue, with a downwardly projecting K/I+ deep blue tubular structure. Ascospores submuriform to muriform, with 5-9 transverse septa and 1-3-longitudinal septa, thin-walled, hyaline, broadly ellipsoid to subfusiform, (20-)25-40(-50) x 11-16 μm. Photobiont cyanobacterial (Nostoc, the cells in short chains). Spot test: all negative. Chemistry: without lichen substances. Note: a widespread holarctic lichen mostly found on soil and amongst mosses in dry grasslands, more rarely on basal parts of trunks. Many records could refer to S. pulvinatum (see notes to that species, to S. imbricatum, and to S. gelatinosum).
Growth form: Squamulose
Substrata: bark, soil, terricolous mosses, and plant debris
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: common
Subalpine belt: extremely common
Oromediterranean belt: common
Montane belt: extremely common
Submediterranean belt: very common
Padanian area: extremely rare
Humid submediterranean belt: very common
Humid mediterranean belt: common
Dry mediterranean belt: rather rare
Predictive model
Herbarium samples
Felix Schumm CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5234)
2001/11/26
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Ampezzo Carnico, 650 m
09.09.2016
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
P.L. Nimis CC BY-SA 4.0
TSB 37971
P.L. Nimis CC BY-SA 4.0
TSB 37971
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Barrage du Tech
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Barrage du Tech
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Le Faouet
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Dineault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Dineault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Dineault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Dineault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Saint Herbot
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Saint Herbot
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Saint Herbot
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Tregarvan
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Fontenay Le Comte
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Fontenay Le Comte
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Fontenay Le Comte
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Portugal: Madeira.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Portugal: Madeira.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Portugal: Madeira.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
Growth form: Squamulose
Substrata: bark, soil, terricolous mosses, and plant debris
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: common
Subalpine belt: extremely common
Oromediterranean belt: common
Montane belt: extremely common
Submediterranean belt: very common
Padanian area: extremely rare
Humid submediterranean belt: very common
Humid mediterranean belt: common
Dry mediterranean belt: rather rare
Predictive model
| Herbarium samples |
Index Fungorum
GBIF
Felix Schumm CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5234)
2001/11/26
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Ampezzo Carnico, 650 m
09.09.2016
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[12319], Griechenland, N-Samos, südlich Avlakia an der Straße nach Vourliotes, 37°48'N, 26°51'E, 163 m; schattige Straßenböschung mit Quercus ilex, Arbutus andrachne u. Styrax-Vegetation. Leg. Schumm, Düll & Maier 03.05.2006, det Schumm 2006.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
P.L. Nimis CC BY-SA 4.0
TSB 37971
P.L. Nimis CC BY-SA 4.0
TSB 37971
Felix Schumm - CC BY-SA 4.0
[16507], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010.
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_S/Text_S/Scytinium_lichenoides.htm
France, 31/3/2011 - Mont d'Or
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Barrage du Tech
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Barrage du Tech
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Le Faouet
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Dineault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Dineault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Dineault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Dineault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Saint Herbot
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Saint Herbot
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Saint Herbot
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Tregarvan
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Fontenay Le Comte
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Fontenay Le Comte
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=342&lang=en
France, Fontenay Le Comte
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Portugal: Madeira.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Portugal: Madeira.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Portugal: Madeira.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.
