Umbilicaria crustulosa (Ach.) Lamy subsp. crustulosa
Bull. Soc. bot. Fr., 25: 386, 1879. Basionym: Gyrophora crustulosa Ach. - Lichenogr. Univ.: 673, 1810.
Synonyms: Gyrophora cirrhosa auct. p.p. non (Hoffm.) Vain.; Gyrophora depressa (Ach.) Röhl.; Gyrophora depressa var. crustulosa (Ach.) Dalla Torre & Sarnth.; Gyrophora spodochroa var. depressa Ach.; Omphalodiscus crustulosus (Ach.) Schol.; Umbilicaria crustulosa (Ach.) Lamy subsp. crustulosa var. crustulosa; Umbilicaria depressa (Ach.) Duby; Umbilicaria spodochroa auct. medioeur. p.p. non Ehrh. ex Hoffm.
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven, TAA (Thor & Nascimbene 2007, Lang 2009, Nascimbene & al. 2022), Lomb (Rivellini 1994, Gheza 2019), Piem (Morisi & Sereno 1995, Isocrono & Falletti 1999, Isocrono & al. 2004, 2006, Morisi 2005, Isocrono & Piervittori 2008), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Isocrono & al. 2008, Favero-Longo & Piervittori 2009), Emil (Fariselli & al. 2020), Lig (Petrak 1965, Brunialti & al. 1999). C - Tosc (Vezda Lich.Rar.Exs. 179!), Sar (Nöske 2000). S - Cal (Puntillo 1996), Si (Ottonello & Isocrono 2004).
Description: Thallus foliose-umbilicate, heteromerous, dorsiventral, monophyllous, sometimes deeply dissected and appearing polyphyllous, orbicular in outline, 3-5(-10) cm wide, rarely wider. 0.2-0.3(-0.5) mm thick. Upper surface pale to dark grey, sometimes with a brown tinge, darker towards the margins, dull, even or slightly radially wrinkled and distinctly areolate around the umbilicus, scabrous and rimose at margin, which are often down-turned and sometimes deeply incised. Lower surface pale grey-white tinged pink, smooth to slightly areolate towards the margins, with numerous cylindrical, simple or branched, concolorous to brown-black rhizinomorphs, which usually lack around the umbilicus; thalloconidia absent. Upper cortex c. 30 µm thick; medulla white; lower cortex c. 40 µm thick. Apothecia common, omphalodisc, sessile to depressed, black, to 5 mm across, with a central sterile button, occasionally appearing subgyrose. Epithecium brown; hymenium colurless, interrupted by bundles of dark-pigmented hyphae; ; paraphyses simple or sparingly branched, septate, the apical cells more or less swollen; hypothecium brown. Asci 8-spored, clavate, thick-walled, with an amyloid apical dome, Umbilicaria-type. Ascospores at first simple and hyaline, then muriform and brown (12-)18-24 x 11-21 µm. Pycnidia numerous in the marginal zone of upper surface, immersed, bottle-shaped, with a dark brown wall and a black ostiole. Conidia bacilliform, c. 3-4 x 0.8-1 µm. Photobiont chlorococcoid. Spot tests: cortex K-, C-, KC-, P-; medulla K-, C+ red, KC+ red, P-. Chemistry: medulla with gyrophoric acid, and small amounts of lecanoric and hiascic acids. Note: a widespread, cool-temperate to arctic-alpine, circumpolar lichen found on exposed, often steeply inclined surfaces of siliceous rocks with some water seepage in upland areas, reaching the nival belt in the Alps; rare in the Apennines due to the scarcity of suitable substrata.
Growth form: Foliose, umbilicate
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
On otherwise dry surfaces with short periods of water seepage after rain
Commonnes-rarity: (info)
Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: rather rare
Montane belt: very rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent
Predictive model
Herbarium samples
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34438)
2002/01/14
Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Felix Schumm – CC BY-SA 4.0
[4262], Schweiz, Flüela Paß südöstlich von Davos, beim O'spiz, auf Granit, ca. 2410 m. Leg. Schäfer-Verwimp & Schumm 27.08.1995, det. Schumm.
Frey E. 1933. Cladoniaceae (unter Ausschluß ded Gattung Cladonia) und Umbilicariaceae. In: Rabenhrst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz. 15: 1-208.
Frey E. 1933. Cladoniaceae (unter Ausschluß ded Gattung Cladonia) und Umbilicariaceae. In: Rabenhrst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz. 15: 1-208.
As U. depressa
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_crustulosa.htm
France, sur gneiss - session AFL 2010 - Hérault
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_crustulosa.htm
France, sur gneiss - session AFL 2010 - Hérault
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_crustulosa.htm
France, sur gneiss - session AFL 2010 - Hérault
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_crustulosa.htm
France, sur gneiss - session AFL 2010 - Hérault
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Corse, Col de Bavella
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Corse, Col de Bavella
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Corse, Col de Bavella
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Corse, Lac Nino, 1800 m
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Corse, Lac Nino, 1800 m
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Pays Basque, Iraty
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Pays Basque, Iraty
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Pays Basque, Iraty
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1240&lang=en
France, Pays Basque, Iraty
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, Villamor (Somiedo), 16-III-2023, en rocas ácidas, junto a Umbilicaria torrefacta y Umbilicaria polyphylla, leg. & det. M. González, MGG-3.
Felix Schumm - CC BY-SA 4.0
Gyrophoric acid
K-, C+ rosa. KC-, P-.
Umkristallisation in heißem bis kochendem G.E. (Glycerin + Eisessig 2:1), iddentifizierbare Kristalle in 30 Minuten bis 24 Stunden
Standardflechte: Umbilicaria crustulosa
Felix Schumm - CC BY-SA 4.0
Gyrophoric acid
K-, C+ rosa. KC-, P-.
Umkristallisation in G.A.W. (Glycerin + 95% Alkohol + Wasser 1:1:1) (Glycerin + Eisessig 2:1), Standardflechte: Umbilicaria crustulosa
Growth form: Foliose, umbilicate
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
On otherwise dry surfaces with short periods of water seepage after rain
Commonnes-rarity: (info)
Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: rather rare
Montane belt: very rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent
Predictive model
Herbarium samples |