Umbilicaria cylindrica (L.) Delise var. cylindrica

in Duby, Bot. Gall., 2: 595, 1830. Basionym: Lichen cylindricus L. - Sp. Pl., 2: 1144, 1753.
Synonyms: Gyrophora cylindrica (L.) Ach.; Gyrophora cylindrica var. nudiuscula (Schaer.) Flot.
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven, TAA (Caniglia & al. 2002, Nascimbene 2003, 2006c, 2008b, Lang 2009, Nascimbene & al. 2022), Lomb (Rivellini 1994, Dalle Vedove & al. 2004, Brackel 2013, Gheza 2019, 2019b), Piem (Morisi & Sereno 1995, Isocrono & Falletti 1999, Isocrono & al. 2003, 2004, 2006, Morisi 2005, Isocrono & Piervittori 2008, Giordani & al. 2014, Favero-Longo & al. 2015, Lich. Ital. Exs. 39: Isocrono & al. 2020), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Piervittori & al. 1998, 2001, 2004, Valcuvia 2000, Isocrono & al. 2008, Favero-Longo & Piervittori 2009, Blisa & al. 2011, Matteucci & al. 2013b, 2015c, Watson 2014), Emil (Dalle Vedove & al. 2002, Fariselli & al. 2020), Lig (Brunialti & al. 1999). C - Tosc, Abr (Gheza & al. 2021), Sar. S - Bas (Potenza & al. 2014), Cal (Puntillo 1996, Potenza & al. 2011).
Description: Thallus foliose-umbilicate, heteromerous, dorsiventral, monophyllous, 3-5(-7) cm wide, 02-0.4 mm thick, the lobe margins with conspicuous but rather sparse, flat, branched, up to 2 mm long black cilia. Upper surface pale to dark grey, sometimes tinged brown, dull, slightly wrinkled or almost even, sometimes with a few folds and ridges around the umbilicus, giving way to a reticulate pattern. Lower surface grey-white, tan or pink, sometimes pink-brown towards the umbilicus and grey-pruinose in the marginal zone, even, smooth, the area around the umbilicus naked, the rest with numerous thin, long, cylindrical, pale, simple rhizinomorphs; thalloconidia absent. Upper cortex 10-20 µm thick; medulla white, 3-layered, the upper and lower layers loose, with an arachnoid plectenchyma, the layer inbetween with a compact plectenchyma; lower cortex 10-20 µm thick, scleroplectenchymatous. Apothecia numerous, black, gyrodisc, markedly stipitate, to 2-5 mm across, convex with few deep gyri. Epithecium dark brown, up to 20 µm high; hymenium colourless, c. 70 µm high; paraphyses usually simple, up to 2.5 µm thick, the apical cells hardly swollen; hypothecium dark. Asci 8-spored, clavate, thick-walled, with an amyloid apical dome, Umbilicaria-type. Ascospores 1-celled, hyaline, ellipsoid, 7-15 x 3-9 µm. Pycnidia numerous, immersed, bottle-shaped, with a brown wall, and a dark brown ostiole. Conidia c. 3-4 x 0.8 µm. Photobiont chlorococcoid. Spot tests: medulla K- or K+ yellow turning red, C-, KC- or KC+ faintly red, P- or P+ orange. Chemistry: medulla with variable amounts of norstictic acid and sometimes with very low amounts of gyrophoric and lecanoric acids.
Note: an ecologically wide-ranging, cool-temperate to arctic-alpine, circumpolar lichen found on wind-exposed boulders with a short snow-covering period, often on or near the top, with optimum above treeline, reaching the nival belt in the Alps; common only in the Alps, becoming much rarer southwards, also for the scarcity of suitable substrata, up to the mountains of Calabria.
Growth form: Foliose, umbilicate
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: common
Subalpine belt: very common
Oromediterranean belt: rare
Montane belt: rather rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/
Austria, Zirbitz



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (8311)
2001/12/05
apothecium



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (8311)
2001/12/05



Felix Schumm – CC BY-SA 4.0
[1531], Österreich, Großglockner Hochalpenstraße, auf Silikatgestein, ca. 1800 m. Leg. Schumm 05.08.1969, det. Schumm 1969



Felix Schumm – CC BY-SA 4.0
[1531], Österreich, Großglockner Hochalpenstraße, auf Silikatgestein, ca. 1800 m. Leg. Schumm 05.08.1969, det. Schumm 1969



Felix Schumm – CC BY-SA 4.0
[1531], Österreich, Großglockner Hochalpenstraße, auf Silikatgestein, ca. 1800 m. Leg. Schumm 05.08.1969, det. Schumm 1969



Felix Schumm – CC BY-SA 4.0
[1531], Österreich, Großglockner Hochalpenstraße, auf Silikatgestein, ca. 1800 m. Leg. Schumm 05.08.1969, det. Schumm 1969



Felix Schumm – CC BY-SA 4.0
[1531], Österreich, Großglockner Hochalpenstraße, auf Silikatgestein, ca. 1800 m. Leg. Schumm 05.08.1969, det. Schumm 1969



Felix Schumm – CC BY-SA 4.0
[1531], Österreich, Großglockner Hochalpenstraße, auf Silikatgestein, ca. 1800 m. Leg. Schumm 05.08.1969, det. Schumm 1969



Curtis Randall Björk, - CC BY-SA 4.0
Sugarbowl-Grizzly Den Provincial Park, British Columbia, Canada
August 2017
as v. tilesii


Helena Rodriguez
Spain, Galicia, Peña Trevinca
06.2019


Frey E. 1933. Cladoniaceae (unter Ausschluß ded Gattung Cladonia) und Umbilicariaceae. In: Rabenhrst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz. 15: 1-208.


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.


Giovanna Potenza
Italy, Basilicata, M. Volturino


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_cylindrica.htm
France, https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_crustulosa.htm


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_cylindrica.htm
France, https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_crustulosa.htm


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
Italy, Val d'Aosta


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
Italy, Val d'Aosta


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
Italy, Val d'Aosta


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
Italy, Val d'Aosta


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
Italy, Val d'Aosta


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
France, Mercantour


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
France, Tarentaise


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
France, Tarentaise


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
France, Tarentaise


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
France, Tarentaise


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
France, Tarentaise


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=549&lang=en
France, Tarentaise