Umbilicaria freyi Codogno, Poelt & Puntillo

Plant Syst. Evol., 165: 63, 1989.
Synonyms: Umbilicaria grisea f. subpapyria Frey; Umbilicaria hirsuta var. pyrenaica Frey
Distribution: N - VA (Piervittori & Isocrono 1999), Emil (Fariselli & al. 2020). C - Tosc, Sar (Narui & al. 1996). S - Cal (Codogno 1995, Puntillo 1996, Hafellner 2021).
Description: Thallus foliose, umbilicate, heteromerous, dorsiventral, attached by a central holdfast, rather thick and rigid, up to 10(-15) cm in diam. Upper surface ochraceous grey to pale grey-brown, dull, finely rimose, the marginal parts and sometimes also the central parts with numerous, up to 1(-2) mm wide, darker brown schizidia which tend to fall off leaving scars exposing the medulla; lower surface ochraceous white to pale pinkish brown, usually distinctly areolate, with or without pale rhizinomorphs, without trabeculae near the central holdfast. Apothecia very rare and incompletely developed, black, gyrodisc. Asci 8-spored, elongate-clavate, thick-walled, the apical dome K/I+ blue, Umbilicaria-type. Ascospores 1-celled, hyaline, ellipsoid, 8-11 x 6-8 μm. Photobiont chlorococcoid. Spot tests: cortex K-, C-, KC-, P-; medulla K_, C+ red, KC+ red, P-. Chemistry: medulla with gyrophoric acid.
Note: on steeply inclined to rain-sheltered surfaces of siliceous rocks, ecologically intermediate between U. grisea and U. deusta, ranging from the Alps to the mountains of Calabria.
Growth form: Foliose, umbilicate
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: rather rare
Oromediterranean belt: rare
Montane belt: rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_freyi.htm
France, Boccadi Vezzu - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Umbilicaria_freyi.htm
France, Boccadi Vezzu - Corse



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11046)
2001/12/05
detail of lower surface with rhizinomorphs



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11083)
2003/03/17


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
France, Corse, Forêt de Tartagine


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
France, Corse, Forêt de Tartagine


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
France, Corse, Forêt de Tartagine


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
France, Corse, Forêt de Tartagine


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
France, Corse, Lumio


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
France, Corse, Lumio


Antonio Valero Risoto - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
Spain, Carnota


Antonio Valero Risoto - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
Spain, Carnota


Antonio Valero Risoto - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
Spain, Carnota


Antonio Valero Risoto - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1237&lang=en
Spain, Carnota