Umbilicaria nylanderiana (Zahlbr.) H. Magn.

Lich. Sel. Scand. Exs.: 252, 1937. Basionym: Gyrophora nylanderiana Zahlbr. - Cat. Lich. Univ, 4: 720, 1927.
Synonyms: Umbilicaria corrugata Nyl.
Distribution: N - TAA (Nascimbene 2008b, Nascimbene & al. 2022), Lomb (Gheza 2019b), Piem (Isocrono & al. 2004, Isocrono & Piervittori 2008), VA (Piervittori & Isocrono 1999, Piervittori & al. 2001, 2004). C - Tosc (Tretiach & al. 2008), Sar. S - Bas (Potenza & al. 2014), Cal (Puntillo 1996), Si (Ottonello & Isocrono 2004, Iacolino & Ottonello 2006).
Description: Thallus foliose-umbilicate, heteromerous, dorsiventral, monophyllous but often deeply dissected and appearing polyphyllous, 2-3(-5) cm wide, attached by a thick, black central holdfast, with often lacerate and perforate, ascending and undulating margins. Upper surface dark brown, sometimes with a thin white necrotic layer over the umbilicus, rugose with even, smooth whenvermiform ridges, towards the margins more or less young, later rugose, especially in central parts. Lower surface sooty black, sometimes slightly paler at margins, smooth, even or dimpled, erhizinate; thalloconidia simple, rarely 1-septate, spherical, (4-)6-9(-11) x (4-)5-6(-11) µm, dark brown, covering the whole lower surface except sometimes the marginal zone. Upper cortex dark brown, 10-20 µm thick; medulla white, 50-150 µm thick; lower cortex 10-20 µm thick. Apothecia very rare, black, gyrodisc. Epithecium brown; hymenium colourless; paraphyses simple or sparingly branched, the apical cells more or less swollen; hypothecium dark. Ascospores 1-celled hyaline, 7-14 x 5-7 µm. Pycnidia mainly marginal on upper surface, immersed, globose, with a dark brown ostiole. Conidia bacilliform. Photobiont chlorococcoid. Spot tests: medulla K-, C+ red, KC+ red, P-. Chemistry: medulla with gyrophoric, lecanoric and umbilicaric acids.
Note: an arctic-alpine, probably circumpolar lichen found on the top of isolated siliceous boulders, most frequent in the Alps above treeline, but reaching south to the mountains of Calabria.
Growth form: Foliose, umbilicate
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by conidia and thalloconidia

Commonnes-rarity: (info)

Alpine belt: rather rare
Subalpine belt: rare
Oromediterranean belt: very rare
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples
Index Fungorum GBIF


Felix Schumm – CC BY-SA 4.0
[19025], Deutschland, Niedersachsen, Nationalpark Harz, Gebirgskamm 'Auf dem, Acker', Kanapeeklippe, auf Quarzit. MTB: 4228/3. Leg. Hans Ullrich 12.06.1960. Dubl. aus NP Harz Nr. 2145.



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (11049)
2001/12/05
detail of upper surface with apothecia



Felix Schumm – CC BY-SA 4.0
[19025], Deutschland, Niedersachsen, Nationalpark Harz, Gebirgskamm 'Auf dem, Acker', Kanapeeklippe, auf Quarzit. MTB: 4228/3. Leg. Hans Ullrich 12.06.1960. Dubl. aus NP Harz Nr. 2145.



Felix Schumm – CC BY-SA 4.0
[19025], Deutschland, Niedersachsen, Nationalpark Harz, Gebirgskamm 'Auf dem, Acker', Kanapeeklippe, auf Quarzit. MTB: 4228/3. Leg. Hans Ullrich 12.06.1960. Dubl. aus NP Harz Nr. 2145.



Felix Schumm – CC BY-SA 4.0
[19025], Deutschland, Niedersachsen, Nationalpark Harz, Gebirgskamm 'Auf dem, Acker', Kanapeeklippe, auf Quarzit. MTB: 4228/3. Leg. Hans Ullrich 12.06.1960. Dubl. aus NP Harz Nr. 2145.



Felix Schumm – CC BY-SA 4.0
[19025], Deutschland, Niedersachsen, Nationalpark Harz, Gebirgskamm 'Auf dem, Acker', Kanapeeklippe, auf Quarzit. MTB: 4228/3. Leg. Hans Ullrich 12.06.1960. Dubl. aus NP Harz Nr. 2145.



Felix Schumm – CC BY-SA 4.0
[19025], Deutschland, Niedersachsen, Nationalpark Harz, Gebirgskamm 'Auf dem, Acker', Kanapeeklippe, auf Quarzit. MTB: 4228/3. Leg. Hans Ullrich 12.06.1960. Dubl. aus NP Harz Nr. 2145.



Felix Schumm – CC BY-SA 4.0
[19025], Deutschland, Niedersachsen, Nationalpark Harz, Gebirgskamm 'Auf dem, Acker', Kanapeeklippe, auf Quarzit. MTB: 4228/3. Leg. Hans Ullrich 12.06.1960. Dubl. aus NP Harz Nr. 2145.



Felix Schumm - CC BY-SA 4.0
[19025], Deutschland, Niedersachsen, Nationalpark Harz, Gebirgskamm 'Auf dem, Acker', Kanapeeklippe, auf Quarzit. MTB: 4228/3. Leg. Hans Ullrich 12.06.1960. Dubl. aus NP Harz Nr. 2145.


Frey E. 1933. Cladoniaceae (unter Ausschluß ded Gattung Cladonia) und Umbilicariaceae. In: Rabenhrst’s Kryptogamenflora von Deutschland, Österreich und der Schweiz. 15: 1-208.
As U. corrugata


Giovanna Potenza
Italy, Basilicata, M. Volturino


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_U/Textes_U/Umbilicaria_nylanderiana.htm
France, session AFL en Haute- Ubaye - Alpes-de-Haute-Provence