Verrucaria alpigena Breuss

Sauteria, 15: 122, 2008.
Synonyms: Verrucaria alpina (Arnold) Breuss non (Bagl. & Carestia) Stizenb.; Verrucaria alpina Breuss; Verrucaria muralis var. alpina Arnold
Distribution: N - TAA (De Benetti & Caniglia 1993).
Description: Thallus crustose, endo- to thinly episubstratic, whitish to pale greyish brown, often with medium brown small flecks, smooth to rimose, to 0.15 mm thick, sometimes delimited by a poorly developed, medium brown, weakly fimbriate prothallus. Cortex usually unpigmented or with a dilute brown pigment. Perithecia black, sometimes greyish in lower part but not covered by a thalline layer, c. 80-100/cm2, 1/2 to 3/4 immersed, forming 0.25-0.5 mm wide, hemispherical to conical projections, the apex rounded to slightly flattened, the base usually not in pits of the rock. Involucrellum present around the upper half of exciple or reaching to base-level, appressed to exciple; exciple 0.2-0.3(-0.4) mm across, colourless to brownish at base; hamathecium of periphysoids, interascal filaments absent; hymenial gel hemiamyloid, I+ red (I+ blue at very low concentrations of I), K/I+ blue. Asci 8-spored, clavate, I-, fissitunicate, the wall thickened above, with an ocular chamber, dehiscent by extrusion of an endotunica to form a delicate rostrum, Verrucaria-type. Ascospores 1-celled, hyaline, ellipsoid, (25-)27-35(-37) x (12-)13-16(-18) μm. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: on calciferous rocks in upland areas, reported from the Alps and the Carpathians; originally described as related to V. muralis, differing in the larger spores (see Breuss 2008b), but according to Pykälä & al. (2020) probably related to V. ahtii in the Endocarpon-group.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Poorly known taxon in need of further study

Commonnes-rarity: (info)

Alpine belt: extremely rare
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: extremely rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model