Verrucaria papillosa Ach.

Lichenogr. Univ.: 286, 1810.
Synonyms:
Distribution: N - Frl (TSB 26483), Ven, TAA, Lomb, Lig (Giordani & al. 2025). C - Tosc. S - Camp, Si.
Description: Thallus crustose, thinly episubstratic, often visible as dirty white to grey flecks, when well-developed rimose-areolate, with or without a dark prothallus, the areoles 0.2-1 mm wide, flat to slightly convex. Perithecia black, 0.2-0.5 mm across, 1/-2/3 immersed, the projecting part mostly hemispherical. Involucrellum black, at least dimidiate but often reaching to base-level, strongly diverging from exciple and wider than the perithecium; exciple globose, 0.3-0.6 mm across, the wall 25-50 μm thick, dark brown to black throughout, K+ darker brown; hamathecium of periphysoids measuring 40-60 x 1.5-2.5 µm, interascal filaments absent; hymenial gel hemiamyloid, I+ red (I+ blue at very low concentrations of I), K/I+ blue. Asci 8-spored, clavate, I-, fissitunicate, the wall thickened above, with an ocular chamber, dehiscent by extrusion of an endotunica to form a delicate rostrum, Verrucaria-type, 90-135 x 30-45 µm. Ascospores 1-celled, hyaline, ellipsoid to broadly ellipsoid, (24-)30-36(-40) x 14-24 μm. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: on more or less calciferous rocks in rather humid situations; similar to V. viridula, but a distinct species according to McCarthy (1988), Breuss (2007, 2008) and Roux & Coll. (2025).
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Poorly known taxon in need of further study

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis CC BY-SA 4.0
TSB 26483



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (26483)


Zschacke, H. (1934) Epigloeaceae, Verrucariaceae und Dermatocarpaceae. In: Dr. L. Rabenhorst‘s Kryptogamen-Flora, Band 9, Abt. 1, Teil 1. Akademische Verlagsgesellschaft, Leipzig, 695 pp. - Public Domain



Pier Luigi Nimis
Herbarium: TSB 27853