Gyalolechia luteococcinea (Cl. Roux) Cl. Roux, M. Bertrand & Arup

in Arup & al., Borziana, 4: 45, 2023. Basionym: Caloplaca subochracea auct. var. luteococcinea Cl. Roux - Bull. Soc. Linn. Provence, 54: 120, 2003
Synonyms: Caloplaca africana sensu Clauzade non (Flagey) Clauzade
Distribution: C - Tosc (Arup & al. 2023), Laz (Arup & al. 2023). S - Pugl (Arup & al. 2023), Si (Arup & al. 2023).
Description: Thallus crustose, episubstratic, finely rimose-areolate, bright yellow to yellow-orange, often delimited by a thin, blue-black prothalline line. Apothecia biatorine, at first immersed, then sessile, rounded, 0.2-1.3 mm across, with a rusty red to dark orange-red, at first flat, then convex, epruinose disc, and a somewhat paler, smooth, at first well-visible, finally often excluded proper margin. Epithecium brownish orange, K+ red; hymenium colourless, 40-60(-80) µm high; paraphyses simple or forked in upper part, not anastomosing, c. 1 µm thick at base, the apical cells (2-)3-4.5 µm wide; hypothecium colourless. Asci 8-spored, clavate, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 2-celled, polarilocular, hyaline, ellipsoid, (9-)10-13(-14) x (3-)5-6(-10) µm, the equatorial thickening (“septum”) up to 5 µm. Photobiont chlorococcoid. Spot tests: thallus K+ red; C-, KC-, P-; apothecia K+ red. Chemistry: thallus with fragilin; apothecia with parietin and emodin (major), and citreorosein, emodinal, fallacinal and parietinic acid (minor).
Note: a Mediterranean lichen found on more or less sheltered calcareous rocks, usually not far from the coast but not strictly maritime.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: rare
Dry mediterranean belt: very rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Michel Bertrand
France, Provence, Bouches-du-Rhône, Marseille, île de Riou


Pier Luigi Nimis
Italy, Sicily, Trapani, Erice, old city walls near the Castle
12/2016



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sicily, Trapani, Erice, castle
12/2016
together with G. marmorata (white thallus)



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Sicily, Trapani, Erice, castle
12/2016
together with G. marmorata (white thallus)



Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710.
var. luteococcinea



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Sicily, Trapani, Erice, Historical Center



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710.
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972.
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972.
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972.
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19605], Hispania, Gerona, Rosasa, Cabo Morfeo, ad saxa calcarea. Leg. X. Llimona, 9.09.1976, det. M. Egea. VEZDA: LICHENES SELECTI EXSICCATI NR. 1972
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710
var. luteococcinea


Department of Life Sciences, University of Trieste
Herbarium: TSB (11674)
2001/11/14



P.L. Nimis
Vezda Lich. Sel. Exs.



Felix Schumm - CC BY-SA 4.0
[19604], Hispania, Levant, Penon de Ifach, alt. 150 m, in vicinitate maris, ad saxa calcarea. Leg. Cl. Roux, 25.08.1977, det. G. Clauzade. VEZDA: LICHENES SELECTI EXSICCATI NR. 1710.
var. luteococcinea