Cetraria crispiformis (Räsänen) Makryǐ

The Lichen Flora of Russia, 3: 84, 2022.. Basionym: Cetraria islandica var. crispiformis Räsänen - Ann. bot. Soc. Zool.-Bot. fenn. Vanamo, 18, 1): 19, 1943.
Synonyms: Cetraria islandica subsp. crispiformis (Räsänen) Kärnefelt
Distribution:
Description: Thallus subfoliose to subfruticose, loosely attached, often forming large tufts, consisting of dorsiventral, flattened, elongate, ascending, 1-15(-30) mm wide and 3-10 cm tall, often sparingly branched laciniae with often in-rolled margins without a distinct ledge, bearing short, marginal, pycnidia-bearing spinulose outgrowths. Upper surface dark brown in sun-forms to pale olive-green in shade, matt or somehow glossy, usually pitted and ridged, the basal parts often reddish; lower surface usually paler, with small, maculiform, irregular, white pseudocyphellae spread throughout the surface and the margins. Cortex 2-layered, with an external layer of brownish, thick-walled, paraplectenchymatous cells, and an inner layer of periclinally arranged hyphae; medulla white. Apothecia extremely rare, lecanorine, mostly subterminal, with a dark brown disc and an often crenulate thalline margin. Ascospores not known. Pycnidia immersed at the top of the marginal spinules, often empty. Conidia simple, elongate, slightly swollen in the middle, 6-7 x c. 1 μm. Photobiont chlorococcoid. Spot tests: cortex K-, C-, KC-, P-; medulla K- or K+ yellow-orange, C-, KC-, P+ yellow to orange or P-, UV-. Chemistry: medulla with fumarprotocetraric, lichesterinic and protolichersterinic acids.
Note: on acid soil in the understory of subalpine coniferous forests and alpine heaths. An arctic-alpine species, most common in Scandinavia and in the British Isles, while from continental Europe there are only a few records; rare in the Alps (outside Italian territory, see Nimis & al. 2018), but perhaps overlooked or not distinguished fromC. islandica and/or C. ericetorum. To be looked for in the Italian Alps.
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by thallus fragmentation

pH of the substrata:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Curtis Randall Björk – CC BY-SA 4.0
Northwest Territories, near MacKay Lake, 2011-08-02



Curtis Randall Björk – CC BY-SA 4.0
Northwest Territories, near MacKay Lake, 2011-08-02



Pier Luigi Nimis - CC BY-SA 4.0
TSB 29500