Cladonia caespiticia (Pers.) Flörke

Clad. Comm.: 8, 1828. Basionym: Baeomyces caespiticius Pers. - Ann. Bot. (Usteri), 7: 155, 1793 (“1794”).
Synonyms: Cladonia agariciformis (Wulfen) Arnold
Distribution: N - VG, Frl (Burgaz & al. 2020), Ven, TAA, Lomb (Rivellini & Valcuvia 1996, Zocchi & al. 1997, Stofer 2006, Abramini & al. 2008, Furlanetto 2010, Gheza 2018, 2019, Gheza & al. 2020, 2022), Piem (Isocrono & al. 2004, Gheza 2018, 2020, Gheza & Nascimbene 2024), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Valcuvia & al. 2000b), Emil (Nimis & al. 1996, Fariselli & al. 2020), Lig (Putortì & al. 1999b, Giordani & Incerti 2008, Gheza & al. 2020). C - Tosc (Loppi & al. 1997b, 1999a, Putortì & al. 1999, Loppi & Putortì 2001, Brunialti & al. 2012b), Laz (Stofer 2006), Sar (HAL-7276). S - Camp, Pugl, Cal (Stofer 2006), Si.
Description: Primary thallus squamulose, the squamules persistent, 1-10 mm long, 2-5 mm wide, strongly divided and irregularly incised, ascending, mostly forming dense, low cushions, pale grey-green to brownish green above, white and very finely arachnoid beneath. Podetia very small, 1-5 mm tall, 0.4-1.5 mm thick, partly ecorticate, rarely with a few squamules, cupless, almost translucent when wet (no photobiont!). Apothecia peltate, sessile on squamules or terminal on the podetia, single or clustered, pale to dark brown, more or less translucent when wet, convex. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia brown, semi-immersed on the upper surface of the squamules, ovoid to conical, constricted at base, with a colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K- or K+ yellowish slowly turning brown, C-, KC-, P+ rapidly red, UV-. Chemistry: fumarprotocetraric acid.
Note: a cool-temperate to southern boreal-montane, circumpolar lichen found on mineral, generally sandy-clay soil, occasionally on rotting wood and at the base of ancient trunks in sheltered situations. The records from Southern Italy (Nimis 1993: 225) need reconfirmation.
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: extremely rare
Oromediterranean belt: extremely rare
Montane belt: very rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Helena Rodriguez
Spain, Galicia



Felix Schumm – CC BY-SA 4.0
[17993], Germany, Baden-Württemberg, Kreis Göppingen, nordöstlich von Baiereck im Brillergehren, rasig auf Baumrinde, 48.75909° N, 9.52401° E, 450 m, TK 7224. Leg. Schumm 11.05.2012, det. Schumm 2012.



Felix Schumm – CC BY-SA 4.0
[17993], Germany, Baden-Württemberg, Kreis Göppingen, nordöstlich von Baiereck im Brillergehren, rasig auf Baumrinde, 48.75909° N, 9.52401° E, 450 m, TK 7224. Leg. Schumm 11.05.2012, det. Schumm 2012.



Felix Schumm – CC BY-SA 4.0
[17993], Germany, Baden-Württemberg, Kreis Göppingen, nordöstlich von Baiereck im Brillergehren, rasig auf Baumrinde, 48.75909° N, 9.52401° E, 450 m, TK 7224. Leg. Schumm 11.05.2012, det. Schumm 2012.



Felix Schumm – CC BY-SA 4.0
[17993], Germany, Baden-Württemberg, Kreis Göppingen, nordöstlich von Baiereck im Brillergehren, rasig auf Baumrinde, 48.75909° N, 9.52401° E, 450 m, TK 7224. Leg. Schumm 11.05.2012, det. Schumm 2012.


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5670)
2001/12/04


Nere Sorrentino
Spain, Galicia, Pontevedra


Leif Stridvall
Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/


Nere Sorrentino
Spain, Galicia, Pontevedra



Pier Luigi Nimis - CC BY-SA 4.0
TSB 42110



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41489



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41489



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41489


Einar Timdal - Source: http://nhm2.uio.no/lav/web/index.html - CC BY-NC


Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Lombardia, Brescia, Breno, 1600 m
30/08/2019


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.


Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Central Europe; Germany: Hesse.


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, La Sierra (Corvera-Asturias), 22-VIII-2023, en un talud herboso, bajo Castanea sativa, leg. M. González, det. Ana Rosa Burgaz, MGG-42. (con ácido fumarprotocetrárico).


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, La Sierra (Corvera-Asturias), 22-VIII-2023, en un talud herboso, bajo Castanea sativa, leg. M. González, det. Ana Rosa Burgaz, MGG-42. (con ácido fumarprotocetrárico).


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, La Sierra (Corvera-Asturias), 22-VIII-2023, en un talud herboso, bajo Castanea sativa, leg. M. González, det. Ana Rosa Burgaz, MGG-42. (con ácido fumarprotocetrárico).


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, La Sierra (Corvera-Asturias), 22-VIII-2023, en un talud herboso, bajo Castanea sativa, leg. M. González, det. Ana Rosa Burgaz, MGG-42. (con ácido fumarprotocetrárico).


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, La Sierra (Corvera-Asturias), 22-VIII-2023, en un talud herboso, bajo Castanea sativa, leg. M. González, det. Ana Rosa Burgaz, MGG-42. (con ácido fumarprotocetrárico).


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, La Sierra (Corvera-Asturias), 22-VIII-2023, en un talud herboso, bajo Castanea sativa, leg. M. González, det. Ana Rosa Burgaz, MGG-42. (con ácido fumarprotocetrárico).


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, La Sierra (Corvera-Asturias), 22-VIII-2023, en un talud herboso, bajo Castanea sativa, leg. M. González, det. Ana Rosa Burgaz, MGG-42. (con ácido fumarprotocetrárico).


Marta González Garcia - Centro de Estudios Micologicos Asturianos
Spain, La Sierra (Corvera-Asturias), 22-VIII-2023, en un talud herboso, bajo Castanea sativa, leg. M. González, det. Ana Rosa Burgaz, MGG-42. (con ácido fumarprotocetrárico).