Lempholemma intricatum (Arnold) Zahlbr.

Lempholemma intricatum (Arnold) Zahlbr.

Cat. Lich. Univ., 3: 23, 1924. Basionym: Omphalaria intricata Arnold - Flora, 52: 254, 1869.

Synonyms: Leciophysma fennicum Räsänen; Lempholemma fennicum (Räsänen) Degel.; Synalissa intricata (Arnold) Nyl.

Distribution: N - Frl (Henssen & Tretiach 1995), Ven (Nascimbene & al. 2021), TAA (Nascimbene & al. 2022).

Description: Thallus small-fruticose, filamentous, homoiomerous, gelatinous when wet, dark brown to almost black, sometimes grey-pruinose, consisting of up to 0.2 mm thick and up to 5 mm long, furcate, adpressed, entangled, longitudinally weakly wrinkled branches, forming up to 3 cm wide, small carpets or loose, flattened cushions. Apothecia (pycnoascocarps) rare, laminal, developing from pycnidia, globose, semi-immersed, up to 0.2 mm across, with a brown, finally convex disc. Proper exciple very thin, of parallel hyphae; epithecium brownish; hymenium and hypothecium colourless, the hymenial gel I+ reddish, K/I+ blue; paraphyses mostly simple, not apically thickened, coherent. Asci 8-spored, prototunicate, cylindrical, without a distinct apical apparatus. Ascospores 1-celled, hyaline, subglobose to broadly ellipsoid, 10-14 x 8-11 μm, with a gelatinous perispore which dissolves in K. Pycnidia dark, immersed, pyriform. Conidia acrogenous, bacilliform or fusiform, 4-5 x 1-1.5 μm. Photobiont cyanobacterial (Nostoc, the individual cells 4-5 μm wide, arranged in short chains). Spot tests: all negative. Chemistry: without lichen substances.

Note: on steeply inclined surfaces of calcareous or basic siliceous rocks in seepage tracks, mostly in humid areas, perhaps more widespread in the Alps.

Growth form: Fruticose

Substrata: rocks

Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)

Reproductive strategy: mainly sexual

Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

On otherwise dry surfaces with short periods of water seepage after rain

Commonnes-rarity: (info)

Alpine belt: absent

Subalpine belt: extremely rare

Oromediterranean belt: absent

Montane belt: extremely rare

Submediterranean belt: extremely rare

Padanian area: absent

Humid submediterranean belt: absent

Humid mediterranean belt: absent

Dry mediterranean belt: absent

pH of the substrata:

1

2

3

4

5

(info)

Solar irradiation:

1

2

3

4

5

(info)

Aridity:

1

2

3

4

5

(info)

Eutrophication:

1

2

3

4

5

(info)

Poleotolerance:

0

1

2

3

(info)

Altitudinal distribution:

1

2

3

4

5

6

(info)

Predictive model

Herbarium samples

P.L. Nimis CC BY-SA 4.0

TSB 24284

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste

Herbarium: TSB (24284)
2001/11/26

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste

Herbarium: TSB (31377)
2001/12/08

Felix Schumm; Owner: CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999

Felix Schumm; Owner: CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999

Felix Schumm; Owner: CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999

Felix Schumm; Owner: CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999

Felix Schumm; Owner: CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste

Herbarium: TSB (24284)
2001/11/26

Felix Schumm - CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999.

Felix Schumm - CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999.

Felix Schumm - CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999.

Felix Schumm - CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999.

Felix Schumm - CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999.

Felix Schumm - CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999.

Felix Schumm - CC BY-SA 4.0

[TSB_31377], Italy, Friuli, Carnic Alps, Passo del Pura, Southern slopes of Mt. Nauleni, c. 2100 m, on limestone. Leg. M. Tretiach, 27.07.1999.

Pier Luigi Nimis - CC BY-SA 4.0

TSB 24284

Pier Luigi Nimis - CC BY-SA 4.0

TSB 24284

Pier Luigi Nimis - CC BY-SA 4.0

TSB 24284

Pier Luigi Nimis - CC BY-SA 4.0

TSB 24284

Pier Luigi Nimis - CC BY-SA 4.0

TSB 24284

Pier Luigi Nimis - CC BY-SA 4.0

TSB 24284

Pier Luigi Nimis - CC BY-SA 4.0

TSB 24284

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