Cladonia squamosa var. subsquamosa (Leight.) Vain.

Meddeland. Soc. Fauna Fl. Fenn., 6: 13, 1881. Basionym: Cladonia delicata var. subsquamosa Nyl. ex Leight. - Lich. Fl. Gr. Brit.: 59, 1871.
Synonyms: Cladonia squamosa var. allosquamosa Hennipman; Cladonia subsquamosa (Leight.) Cromb. non Kremp.
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven, Lomb (Rivellini & Valcuvia 1996), Piem (Isocrono & al. 2003, 2004, 2006, Isocrono & Piervittori 2008), VA (Piervittori & Isocrono 1997, 1999, Piervittori & al. 2001, 2004), Lig (Brunialti & al. 1999, Watson 2014, Gheza & al. 2020). C - Tosc (Tretiach & Nimis 1994), Marc (Nimis & Tretiach 1999), Umb (Ravera 2000, Ravera & al. 2006). S - Cal (Puntillo 1996), Si (Nimis & al. 1994).
Description: Primary thallus squamulose, persistent or rarely disappearing, the squamules indented, 2-10 mm long, grey-green above, white beneath, esorediate but sometimes granulose on the lower surface. Podetia extremely variable in shape and size, hollow inside, greenish grey to brown, not melanotic at base, 3-7(-14) cm tall, 0.5-2(-5) mm thick, simple or sparingly branched with open axils, usually expanded in upper parts to form gaping funnels which may proliferate radially, the surface often largely ecorticate exposing the stereome, densely covered with microsquamules, squamules and granules, esorediate. Apothecia infrequent, brown, convex, located at the top of podetia or at the margin of funnels, often clustered. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed on podetial tips, rarely on basal squamules, with a red or colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K+ yellow, C-, KC-, P+ yellow, UV-. Chemistry: thamnolic and barbatic acids, the former mostly present in fertile specimens, and unknown terpenoids.
Note: more hygrophytic than the typical variety, and more bound to higher altitudes, this chemical variety has been often considered as an independent species. According to Aptroot (in litt.) the type material belongs to C. ramulosa.
Growth form: Fruticose

Substrata: lignum, soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: rather rare
Oromediterranean belt: absent
Montane belt: rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples
Index Fungorum GBIF


Felix Schumm – CC BY-SA 4.0
[14149], Portugal, Azoren, Terceira, im Gebiet Terra Brava südlich Algualvar, 38°44.738' N, 27°11.880' W, ca. 650 m. Leg. Aptroot & Schumm 15.07.2008.



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10717)
2001/12/04



Felix Schumm – CC BY-SA 4.0
[14149], Portugal, Azoren, Terceira, im Gebiet Terra Brava südlich Algualvar, 38°44.738' N, 27°11.880' W, ca. 650 m. Leg. Aptroot & Schumm 15.07.2008.



Felix Schumm – CC BY-SA 4.0
[14149], Portugal, Azoren, Terceira, im Gebiet Terra Brava südlich Algualvar, 38°44.738' N, 27°11.880' W, ca. 650 m. Leg. Aptroot & Schumm 15.07.2008.



Felix Schumm – CC BY-SA 4.0
[14149], Portugal, Azoren, Terceira, im Gebiet Terra Brava südlich Algualvar, 38°44.738' N, 27°11.880' W, ca. 650 m. Leg. Aptroot & Schumm 15.07.2008.



Felix Schumm – CC BY-SA 4.0
[14149], Portugal, Azoren, Terceira, im Gebiet Terra Brava südlich Algualvar, 38°44.738' N, 27°11.880' W, ca. 650 m. Leg. Aptroot & Schumm 15.07.2008.



Felix Schumm – CC BY-SA 4.0
[14149], Portugal, Azoren, Terceira, im Gebiet Terra Brava südlich Algualvar, 38°44.738' N, 27°11.880' W, ca. 650 m. Leg. Aptroot & Schumm 15.07.2008.,
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UV-Langwelle. 1: Start 4: Position von Norstictinsäure 7: Position von Atranorin th: Thamnolsäure


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Plougonven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Plougonven


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Pointe de Rostudel


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Huelgoat
pycnidiate form


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Huelgoat


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Huelgoat


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Huelgoat


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Huelgoat


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Landevennec


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Menez, Kador, Crozon


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Menez, Kador, Crozon


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=737&lang=en
France, Cap de la Chèvre


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1389&lang=en
France, Lothey, lignicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1389&lang=en
France, Lothey, lignicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1389&lang=en
France, Lothey, lignicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1389&lang=en
France, Lothey, lignicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1389&lang=en
France, Lothey, lignicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1389&lang=en
France, Lothey, lignicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1389&lang=en
France, Lothey, lignicolous


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1361&lang=en
France, Cap de la Chèvre
form with brown interior of podetia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1361&lang=en
France, Cap de la Chèvre
form with brown interior of podetia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1361&lang=en
France, Cap de la Chèvre
form with brown interior of podetia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1361&lang=en
France, Cragou
form with brown interior of podetia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1361&lang=en
France, Cragou
form with brown interior of podetia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1361&lang=en
France, Cragou
form with brown interior of podetia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1361&lang=en
France, Pointe du Guern
form with brown interior of podetia


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=1361&lang=en
France, Pointe du Guern
form with brown interior of podetia