Clavascidium lacinulatum (Ach.) M. Prieto

in Prieto & al., Am. J. Bot., 99: 28, 2012. Basionym: Endocarpon hepaticum var. lacinulatum Ach. - Lichenogr. Univ.: 299, 1810.
Synonyms: Catapyrenium lacinulatum (Ach.) Breuss; Dermatocarpon trapeziforme auct. non (J. König) Trevis.; Placidium lacinulatum (Ach.) Breuss
Distribution: N - Ven, TAA. C - Sar. S - Pugl, Cal (Puntillo 1996).
Description: Thallus squamulose, pale to dark brown, dull, non- or very slightly pruinose. Squamules , scattered to contiguous, rarely overlapping, 3-8 mm broad, 0.3-0.5 mm thick, at first rounded, then crenulate, flattened to weakly convex, smooth, with an undulate-crisped edge. Lower surface pale brown, sometimes darkening, attached by several, simple to weakly dichotomously branched, robust, 100-300 μm thick, several mm long, pale to brown rhizines, and by colourless, 3-5 μm thick rhizohyphae. Upper cortex paraplectenchymatous 40-80 µm thick; medulla of isodiametrical, thick-walled cells, well-delimited from lower cortex; lower cortex poorly developed, the cells not arranged in vertical rows. Perithecia frequent, laminal, immersed in the squamules, globose to broadly pyriform, up to 0.65 mm across, without involucrellum. Exciple colourless to pale brown throughout; hymenial gel hemiamyloid; hamathecium of periphyses measuring 25-35 x 2.5-3.5 µm, interascal filaments absent. Asci 8-spored, cylindrical, thin-walled, I-, without a tholus, 65-80 x 12-17 µm. Ascospores 1-celled, hyaline, ellipsoid, thin-walled, uniseriately arranged at least in young asci, 12-15(-17) x 5.5-7.5 μm. Pycnidia laminal, immersed, black, to 0.5 mm in diam., Dermatocarpon-type. Conidia oblong-ellipsoid, 3-4(-5) x 1.3-2 μm. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a mainly Mediterranean-Atlantic to mild-temperate terricolous species found on loess and calciferous ground, most frequent in dry grasslands at relatively low elevations.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: rare
Dry mediterranean belt: rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb.Glowacki
2003/04/28



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb.Glowacki
2003/04/28



Pier Luigi Nimis - CC BY-SA 4.0
TSB 41278



P.L. Nimis CC BY-SA 04
GZU Herb.Glowacki


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_P/Text_P_2/Placidium_lacinulatum.htm
France, 4/10/2010 - Saint Florent la Strutta - Corse


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue, dunes


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Michel David - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, La Palue


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=741&lang=en
France, Pointe de Pen-Hir