Description: Thallus squamulose, cream-coloured to dark brown, the squamules 2-5 mm broad, 0.23-0.33 mm thick, rounded to lobulate, flattened to slightly convex, smooth, scattered to contiguous, sometimes overlapping, developing on a dark hypothallus. Lower surface at first pale, but often blackening with age, attached by a mat of 2.5-5 μm thick, dark rhizohyphae and by up to 0.3 mm thick, 0.5-1.5 mm long, mostly simple, sparse, dark rhizines. Upper cortex paraplectenchymatous, 37-100 μm thick, of 6-15 μm wide cells, with or without an epinecral layer; medulla prosoplectenchymatous, of elongated cells; lower cortex absent. Perithecia frequent, laminal, pyriform to subglobose, immersed, up to 0.5 mm wide, without involucrellum, with a 20-55 μm thick, carbonaceous, entire exciple. Paraphyses absent, substituted by periphyses. Asci 8-spored, narrowly clavate, the wall I-. Ascospores 1-celled, hyaline, thin-walled, very variable in shape and size, from ellipsoid to club-shaped, (13-)15-20(-23) x (6-)7-9 μm. Pycnidia laminal, immersed, up to 200 μm wide. Conidia bacilliform, (5-)6-8(-9) x 1-1.5 μm. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a mainly Mediterranean lichen growing in dry grasslands on calcareous substrata.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris