Heterodermia obscurata (Nyl.) Trevis.

N. Giorn. Bot. Ital., 1: 114, 1869. Basionym: Physcia speciosa subsp. obscurata Nyl. - Acta Soc. Sci. Fenn., 7: 440, 1863.
Synonyms: Anaptychia hypoleuca auct. p.p. non (Muhl.) A. Massal.; Anaptychia obscurata (Nyl.) Vain.; Anaptychia sorediifera (Müll. Arg.) Du Rietz & Lynge; Heterodermia hypoleuca auct. p.p.; Pseudophyscia hypoleuca auct. p.p.
Distribution: N - Ven (Nascimbene & Caniglia 2000), TAA (B-Leg. H. Sipman nr. 8585), Emil (Fariselli & al. 2020), Lig (Petrak 1965, Castello & al. 1994, Brunialti & Giordani 2000, 2003, Giordani & al. 2001, 2002, Giordani 2006, Giordani & Incerti 2008, Watson 2014). C - Tosc (Loppi & al. 1999a, Laganà & al. 2002, Lorenzini & al. 2003, Loppi & Frati 2006, Brunialti & Frati 2010, Brunialti & al. 2012b, Paoli & al. 2012, Benesperi & al. 2013), Umb (Ravera 2000, Panfili 2000, Ravera & al. 2006), Laz (Bartoli & al. 1997, Ravera 2001, Massari & Ravera 2002, Stofer 2006, Munzi & al. 2007), Mol (Frati & al. 2004, Caporale & al. 2008, Paoli & al. 2015), Sar (Zedda 2002). S - Camp (Nimis & Tretiach 2004, Nascimbene & al. 2010b, Brunialti & al. 2013, Ravera & Brunialti 2013), Cal (Puntillo 1996, Incerti & Nimis 2006).
Description: Thallus foliose, heteromerous, dorsiventral, narrow-lobed, often forming more or less orbicular rosettes, 2-4 (-10) cm in diam., adnate, dichotomously to irregularly lobate. Lobes c. 1-2 mm wide, radiating, sparsely divided, discrete, flat to slightly convex, minutely notched, eciliate. Upper surface grey-white, somewhat shiny, usually epruinose with granular soredia arranged in labriform soralia at the tip of lobes, and often with marginal cilia. Upper cortex prosoplectenchymatous; medulla white in upper part, orange-brown in lower part; lower cortex absent. Lower surface felted-arachnoid, rust-colored to orange-brown, the pigment reacting K+ purple, with sparse, black, marginal, simple to squarrosely branched rhizines. Apothecia rare, lecanorine, 1-5 mm across, with a blackish brown disc and an often sorediate thlline margin. Asci 8-spored, cylindrical to subclavate, Lecanora- type. Ascospores l-septate, brown, ellipsoid, 29-35 x 15-19 µm, Pachysporaria-type. Photobiont chlorococcoid. Spot tests: cortex K+ yellow, C-, KC-, P+ yellow; medulla K-, C-, KC-, P-, the pigmented parts K+ purple. chemistry: cortex with atranorin and chloroatranorin; medulla with atranorin, zeorin, an unidentified terpene and an orange brown pigment in lower part.
Note: a mild-temperate species found on more or less isolated trees, occasionally on epilithic mosses; mainly Tyrrhenian, and also found in urban areas (e.g. at La Spezia). See also note on Polyblastidium subneglectum. The species will be probably soon transferred to another genus (see note on Heterodermia).
Growth form: Foliose, narrow lobed
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: rather rare
Humid mediterranean belt: rather common
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm - CC BY-SA 4.0
[15470], La Reunion, Ilet à Vidot im Cirque de Salazie (Pcknickplatz); 21,06975°S, 55,51416°E, 936 m. Leg. Schumm & Frahm 13.09.2009, det. Schumm 2013



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (16354)
2008.03.06



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (23341)
2008.03.06



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (23341)
2008.03.06



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20257)
2008.03.06


A. Aptroot; Owner: A. Aptroot


A. Aptroot; Owner: A. Aptroot



Felix Schumm - CC BY-SA 4.0
[15470], La Reunion, Ilet à Vidot im Cirque de Salazie (Pcknickplatz); 21,06975°S, 55,51416°E, 936 m. Leg. Schumm & Frahm 13.09.2009, det. Schumm 2013



Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3



Felix Schumm - CC BY-SA 4.0
[15470], La Reunion, Ilet à Vidot im Cirque de Salazie (Pcknickplatz); 21,06975°S, 55,51416°E, 936 m. Leg. Schumm & Frahm 13.09.2009, det. Schumm 2013



Felix Schumm - CC BY-SA 4.0
[KK19036], Australien, New South Wales, Royal National Park südlich von Sydney. Bola Creek, östlich von Waterfall. In einem temperierten Regenwald. 250 m, 34°08’ S, 151°02’ E. Leg. K. & A. Kalb, 02.08.1988, det. K. Kalb.



Felix Schumm - CC BY-SA 4.0
[KK19036], Australien, New South Wales, Royal National Park südlich von Sydney. Bola Creek, östlich von Waterfall. In einem temperierten Regenwald. 250 m, 34°08’ S, 151°02’ E. Leg. K. & A. Kalb, 02.08.1988, det. K. Kalb.



Felix Schumm - CC BY-SA 4.0
[KK19036], Australien, New South Wales, Royal National Park südlich von Sydney. Bola Creek, östlich von Waterfall. In einem temperierten Regenwald. 250 m, 34°08’ S, 151°02’ E. Leg. K. & A. Kalb, 02.08.1988, det. K. Kalb.



Felix Schumm - CC BY-SA 4.0
[KK19036], Australien, New South Wales, Royal National Park südlich von Sydney. Bola Creek, östlich von Waterfall. In einem temperierten Regenwald. 250 m, 34°08’ S, 151°02’ E. Leg. K. & A. Kalb, 02.08.1988, det. K. Kalb.



Felix Schumm - CC BY-SA 4.0
[KK19036], Australien, New South Wales, Royal National Park südlich von Sydney. Bola Creek, östlich von Waterfall. In einem temperierten Regenwald. 250 m, 34°08’ S, 151°02’ E. Leg. K. & A. Kalb, 02.08.1988, det. K. Kalb.



Felix Schumm - CC BY-SA 4.0
[KK19036], Australien, New South Wales, Royal National Park südlich von Sydney. Bola Creek, östlich von Waterfall. In einem temperierten Regenwald. 250 m, 34°08’ S, 151°02’ E. Leg. K. & A. Kalb, 02.08.1988, det. K. Kalb.



Felix Schumm - CC BY-SA 4.0
[KK19036], Australien, New South Wales, Royal National Park südlich von Sydney. Bola Creek, östlich von Waterfall. In einem temperierten Regenwald. 250 m, 34°08’ S, 151°02’ E. Leg. K. & A. Kalb, 02.08.1988, det. K. Kalb.



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20257)
2008.03.06



Pier Luigi Nimis
Italy
TSB 41750



P.L. Nimis CC BY-SA 04
TSB 9319



P.L. Nimis CC BY-SA 04
TSB 9319



P.L. Nimis CC BY-SA 04
TSB 16354


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_H/Texte_H2/Heterodermia_obscurata.htm
France, Casalabria, Laronu, alt. 200m - Corse - (20) - sur écorce de Quercus suber
29/3/2014
subsp. longispora


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_H/Texte_H2/Heterodermia_obscurata.htm
France, Casalabria, Laronu, alt. 200m - Corse - (20) - sur écorce de Quercus suber
29/3/2014
subsp. longispora


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_H/Texte_H2/Heterodermia_obscurata.htm
France, Casalabria, Laronu, alt. 200m - Corse - (20) - sur écorce de Quercus suber
29/3/2014
subsp. longispora


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_H/Texte_H2/Heterodermia_obscurata.htm
France, Casalabria, Laronu, alt. 200m - Corse - (20) - sur écorce de Quercus suber
29/3/2014
subsp. longispora


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_H/Texte_H2/Heterodermia_obscurata.htm
France, Casalabria, Laronu, alt. 200m - Corse - (20) - sur écorce de Quercus suber
29/3/2014
subsp. longispora