Lecanora cenisia Ach.

Lichenogr. Univ.: 361, 1810.
Synonyms: Lecanora atrynea (Ach.) Nyl.; Lecanora atrynea f. integrella (Schaer.) Jatta; Lecanora atrynea var. melacarpa Nyl.; Lecanora cenisia var. atrynea (Ach.) H. Magn.; Lecanora saligna f. saxicola B. de Lesd.; Lecanora subfusca var. atrynea Ach.; Lecanora transcendens (Nyl.) Arnold; Patellaria cenisia var. integrella (Schaer.) Trevis.; Zeora cenisia (Ach.) Flot.
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven (Nascimbene 2005c, 2008), TAA (Caniglia & al. 2002, Nascimbene 2003, 2006c, 2008b), Lomb (De Vita & Valcuvia 2004, Dalle Vedove & al. 2004), Piem (Morisi & Sereno 1995, Isocrono & al. 2004, Morisi 2005, Favero-Longo & al. 2006b, 2015, Isocrono & Ferrarese 2008, Isocrono & Piervittori 2008), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, 1999, Piervittori & al. 2001, 2004, Matteucci & al. 2008c, 2015c, Isocrono & al. 2008, Favero-Longo & Piervittori 2009), Emil (Dalle Vedove & al. 2002, Tretiach & al. 2008, Benesperi 2009, Fariselli & al. 2020), Lig (TSB 33415). C - Tosc, Laz (TSB 18677), Sar (Nöske 2000, Neuwirth 2018). S - Camp (Ricciardi & al. 2000), Cal (Puntillo 1996), Si.
Description: Thallus crustose, episubstratic, continuous to verrucose-areolate, whitish grey, pale blue-grey or yellowish white, epruinose. Apothecia lecanorine, strongly constricted at base, 0.5-2(2.5) mm across, with a pale red-brown to blackish brown, mostly flat, sometimes slightly pruinose disc, and a thick, long-persistent but sometimes finally almost excluded, smooth to verruculose thalline margin. Thalline exciple corticate, the medulla with large crystals insoluble in K; proper exciple colourless, lacking crystals; epithecium brown to olive-brown, K+ green, N+ purple, with coarse crystals, the pigment and the crystals dissolving in K; hymenium colourless, 60-90 µm high; paraphyses 1.5-2 µm thick in lower part, the apical cells up to 3 µm wide; hypothecium colourless, not inspersed with oil droplets. Asci 8-spored, clavate, very thin-walled, with a K/I+ blue, tall tholus penetrated by a faintly amyloid apical cushion, the wall K/I-, surrounded by a blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, broadly ellipsoid, 9-17(-19) x (4.5-)6-9 µm, the wall >1 µm thick. Photobiont chlorococcoid. Spot tests: thallus K+ yellow or K+ yellow turning red, C-, KC-, P+ pale yellow to orange. Chemistry: atranorin (major), chloroatranorin (minor), roccellic acid (major or absent), sometimes norstictic acid.
Note: a circumpolar, cool-temperate to arctic-alpine lichen of siliceous rocks, more rarely found also on hard lignum, most frequent in upland areas; common and widespread in the Alps, becoming rarer along the Apennines, south to the mountains of Sicily.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: very common
Subalpine belt: extremely common
Oromediterranean belt: rare
Montane belt: rather rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m.
Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35202)
2002/07/09



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35202)
2002/07/09



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (32850)
2001/12/08
v. melacarpa


Andres Saag; Owner: University of Tartu


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php . Courtesy: A. Stridvall


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php . Courtesy: A. Stridvall



Felix Schumm – CC BY-SA 4.0
[618], Germany, Bayern, Bayerischer Wald, Prolergipfel. Leg. E. Schmid 07.1952, det. J. Poelt.



Felix Schumm – CC BY-SA 4.0
[618], Germany, Bayern, Bayerischer Wald, Prolergipfel. Leg. E. Schmid 07.1952, det. J. Poelt.



Felix Schumm – CC BY-SA 4.0
[618], Germany, Bayern, Bayerischer Wald, Prolergipfel. Leg. E. Schmid 07.1952, det. J. Poelt.
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht, 366 nm UV-Langwelle und Wasser vor Säurebehandlung 1: Start 4: Position von Norstictinsäure 7: Position von Atranorin at: Atranorin, roc: Roccellsäure



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UV-Langwelle. 1: Start 4: Position von Norstictinsäure 7: Position von Atranorin at = Atranorin



P.L. Nimis CC BY-SA 4.0
TSB 37374



P.L. Nimis CC BY-SA 4.0
TSB 38498



Felix Schumm – CC BY-SA 4.0
[14132], Portugal, Azoren, Terceira, beim Hotel Terceira Mar, 38°39.347' N, 27°13.924' W, ca. 8 m. Leg. Aptroot & Schumm 14.07.2008, det. Schumm.



P.L. Nimis CC BY-SA 04
TSB 38498