Lecanora expallens Ach.

Lichenogr. Univ.: 374, 1810.
Synonyms: Lecanora conizaea (Ach.) Nyl. non auct.; Lecanora foehrensis Erichsen; Lecidea soraliata Vain.
Distribution: N - VG (Castello 2002, Martellos & Castello 2004, Castello & Skert 2005), Frl (Bernini & al. 2010), Ven (Nascimbene & al. 2005b, 2006c, 2007), TAA (Nascimbene & al. 2006, 2007b, 2014, Zarabska & al. 2009, Nascimbene 2014, Nascimbene & Marini 2015, Trindade & al. 2021), Lomb (Stofer 2006, Gheza & al. 2022b), Piem (Isocrono & al. 2009, Giordani & Malaspina 2016), VA (Matteucci & al. 2008, Isocrono & al. 2008), Emil (Nimis & al. 1996, Fariselli & al. 2020), Lig (Giordani 2006, Giordani & Incerti 2008). C - Tosc (Loppi & al. 1997b, 1999a, Putortì & Loppi 1999, Benesperi 2006, 2011, Benesperi & al. 2007, 2013, Tretiach & al. 2008, Frati & al. 2008, Brunialti & Frati 2010, Paoli & al. 2012, Nascimbene & al. 2015), Marc (Nimis & Tretiach 1999, Frati & Brunialti 2006), Umb (Nimis & Tretiach 1999, Ravera & al. 2006), Laz (Ravera & al. 1999, Massari & Ravera 2002, 2006c, Stofer 2006, Munzi & al. 2007, Ravera & Genovesi 2008), Abr (Nimis & Tretiach 1999, Stofer 2006, Caporale & al. 2016), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Ravera & al. 2010, Paoli & al. 2015, Caporale & Ravera 2020), Sar (Zedda 2002, Stofer 2006, Rizzi & al. 2011, Cossu 2013). S - Camp (Aprile & al. 2002, 2003b, Nimis & Tretiach 2004, Ravera 2008b, Brunialti & al. 2010, 2013, Garofalo & al. 2010, Ravera & Brunialti 2013, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999, Potenza 2006, Potenza & al. 2010), Cal (Puntillo & Puntillo 2004, Stofer 2006), Si (Grillo & Caniglia 2004, 2006, Falco Scampatelli 2005, Stofer 2006).
Description: Thallus crustose, usually endosubstratic in non-sorediate parts, episubstratic in sorediate parts, thin, yellowish green, at first forming small orbicular patches up to 1-2 cm across, which later later become confluent and may cover extensive areas, often delimited by a whitish prothallus. Fertile specimens sometimes with esorediate, slightly convex, to 0.2 mm wide, ecorticate areoles. Sorediate specimens with diffuse, farinose (20-50 µm in diam.), yellowish green soredia which finally cover almost the whole thallus, giving it a subleprose appearance. Apothecia rare, lecanorine, up to 1.5 mm across, with a dull yellow to pale brown, flat to slightly convex disc and a weakly prominent, often persistent, sorediate thalline margin. Thalline exciple of irregularly entangled hyphae; epithecium pale yellow, with numerous small yellowish crystals soluble in K; hymenium colourless; paraphyses simple to weakly branched and anastomosing, 1.5-2.5 µm thick, with slightly thickened apical cells; hypothecium colourless to pale yellow. Asci 8-spored, clavate, very thin-walled, with a K/I+ blue, tall tholus penetrated by a faintly amyloid apical cushion, the wall K/I-, surrounded by a blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, ellipsoid to broadly ellipsoid, (7.5-)9-13(-17) x (3-)4-6(-7.5) µm. Photobiont chlorococcoid. Spot tests: thallus and soredia K+ pale yellow, C+ deep yellow to orange, P-; UV+ brownish orange (long wave). Chemistry: usnic acid, zeorin, thiophanic acid (major), unidentified xanthones, expallens-unknown.
Note: a mainly temperate species found on acid, generally rough bark, especially abundant on Quercus cerris in open woodlands, sometimes on lignum, widespread and locally common throughout the country below the montane belt.
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: common
Padanian area: rare
Humid submediterranean belt: very common
Humid mediterranean belt: rather rare
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm CC BY-SA 4.0
[10390], Spanien, Kanaren, La Gomera, an der Straße bei Acardece nordöstlich von Arure, 28°08.611' N, 17°18.507' W, 940 m, licht- und windoffen an Phoenix canariensis.
Leg. F. Schumm 19.04.2003 (26).



Domenico Puntillo; Owner: Domenico Puntillo



Felix Schumm – CC BY-SA 4.0
[4206], Germany, Baden-Württemberg, Kreis Ludwigsburg, im Krummbachtal zwischen Büsnau und Gerlingen, Waldrand östl. des Rappenhofes; ca. 430 m, auf Quercus, TK 7220/NW. Leg. et det. Schumm 15.05.1995.



Felix Schumm – CC BY-SA 4.0
[9583], Germany, Baden-Württemberg, Stuttgart, zwischen Stuttgart und Leonberg, an Kastanien an der Schattenalle beim Bärensee, 48°45.368' N, 9°05.505' E, 430 m, TK: 7220. Leg. F. Schumm 15.06.2002, det. F. Schumm, conf. S. Bayerova 2002



Felix Schumm – CC BY-SA 4.0
[9583], Germany, Baden-Württemberg, Stuttgart, zwischen Stuttgart und Leonberg, an Kastanien an der Schattenalle beim Bärensee, 48°45.368' N, 9°05.505' E, 430 m, TK: 7220. Leg. F. Schumm 15.06.2002, det. F. Schumm, conf. S. Bayerova 2002



Felix Schumm – CC BY-SA 4.0
[9583], Germany, Baden-Württemberg, Stuttgart, zwischen Stuttgart und Leonberg, an Kastanien an der Schattenalle beim Bärensee, 48°45.368' N, 9°05.505' E, 430 m, TK: 7220. Leg. F. Schumm 15.06.2002, det. F. Schumm, conf. S. Bayerova 2002



Felix Schumm – CC BY-SA 4.0
[9583], Germany, Baden-Württemberg, Stuttgart, zwischen Stuttgart und Leonberg, an Kastanien an der Schattenalle beim Bärensee, 48°45.368' N, 9°05.505' E, 430 m, TK: 7220. Leg. F. Schumm 15.06.2002, det. F. Schumm, conf. S. Bayerova 2002
38 Lecanora expallens u: usnic acid, z: zeorin, no: norstictic acid



Felix Schumm – CC BY-SA 4.0
[9587], Germany, Baden-Württemberg, zwischen Calw und Althengstett, Straßenallee, auf Acer, TK: 7218. Leg. F. Schumm 24.08.1990, det. F. Schumm 2002, conf. S. Bayerova 2002.



Felix Schumm – CC BY-SA 4.0
[9587], Germany, Baden-Württemberg, zwischen Calw und Althengstett, Straßenallee, auf Acer, TK: 7218. Leg. F. Schumm 24.08.1990, det. F. Schumm 2002, conf. S. Bayerova 2002.



Felix Schumm – CC BY-SA 4.0
[9587], Germany, Baden-Württemberg, zwischen Calw und Althengstett, Straßenallee, auf Acer, TK: 7218. Leg. F. Schumm 24.08.1990, det. F. Schumm 2002, conf. S. Bayerova 2002.



Felix Schumm – CC BY-SA 4.0
[10390], Spanien, Kanaren, La Gomera, an der Straße bei Acardece nordöstlich von Arure, 28°08.611' N, 17°18.507' W, 940 m, licht- und windoffen an Phoenix canariensis. Leg. F. Schumm 19.04.2003 (26).



Felix Schumm – CC BY-SA 4.0
[10390], Spanien, Kanaren, La Gomera, an der Straße bei Acardece nordöstlich von Arure, 28°08.611' N, 17°18.507' W, 940 m, licht- und windoffen an Phoenix canariensis. Leg. F. Schumm 19.04.2003 (26).



Felix Schumm – CC BY-SA 4.0
[10390], Spanien, Kanaren, La Gomera, an der Straße bei Acardece nordöstlich von Arure, 28°08.611' N, 17°18.507' W, 940 m, licht- und windoffen an Phoenix canariensis. Leg. F. Schumm 19.04.2003 (26).



Felix Schumm – CC BY-SA 4.0
[10390], Spanien, Kanaren, La Gomera, an der Straße bei Acardece nordöstlich von Arure, 28°08.611' N, 17°18.507' W, 940 m, licht- und windoffen an Phoenix canariensis. Leg. F. Schumm 19.04.2003 (26).
us: usnic acid, z: zeorin, t: thiophanic acid



Felix Schumm – CC BY-SA 4.0
[10390], Spanien, Kanaren, La Gomera, an der Straße bei Acardece nordöstlich von Arure, 28°08.611' N, 17°18.507' W, 940 m, licht- und windoffen an Phoenix canariensis. Leg. F. Schumm 19.04.2003 (26).



P.L. Nimis CC BY-SA 04
TSB 20747



P.L. Nimis CC BY-SA 04
TSB 36978