Lecanora intumescens (Rebent.) Rabenh.

Deutsche Krypt.-Fl., 2: 334, 1845. Basionym: Parmelia intumescens Rebent. - Prodr. Fl. Neomarch.: 301, 1804.
Synonyms: Lecanora intumescens var. glaucorufa (Mart.) Körb.; Lecanora subfusca var. intumescens (Rebent.) Flot.; Ochrolechia parella var. tumidula (Pers.) Arnold non auct.; Ochrolechia tumidula (Pers.) Arnold non auct.; Patellaria intumescens (Rebent.) Trevis.
Distribution: N - VG (Carvalho 1997), Frl, Ven (Lazzarin 1997, 2000, Nascimbene & Caniglia 2003c, Nascimbene & al. 2007, Nascimbene 2008c), TAA (Nascimbene & Caniglia 2000b, 2002c, Nascimbene & al. 2006e, 2007b, Nimis & al. 2015), Lomb (Zocchi & al. 1997), Piem (Caniglia & al. 1992, Arosio & al. 1998, Isocrono & al. 2004, 2006, 2007), Emil (Nimis & al. 1996, Tretiach & al. 2008, Benesperi 2009, Fariselli & al. 2020), Lig (Lumbsch & al. 1997, Putortì & al. 1999b, Brunialti & Giordani 2003, Giordani 2006, Giordani & Incerti 2008, Watson 2014). C - Tosc (Tretiach & Nimis 1994, Loppi & Putortì 1995b, Paoli & Loppi 2001, Frati & al. 2006b, Loppi & al. 2006, Loppi & Frati 2006, Benesperi & al. 2007, Brunialti & Frati 2010, Benesperi 2011, Nascimbene & al. 2015), Marc (Nimis & Tretiach 1999), Umb (Ravera 1998, Nimis & Tretiach 1999, Ravera & al. 2006), Laz (Massari & Ravera 2002), Abr (Nimis & Tretiach 1999, Caporale & al. 2016, Corona & al. 2016, Gheza & al. 2021), Mol (Nimis & Tretiach 1999, Caporale & al. 2008, Ravera & al. 2010, Genovesi & Ravera 2014), Sar (Cossu 2013). S - Camp (Aprile & al. 2002, Nimis & Tretiach 2004, Brunialti & al. 2010, 2013, Garofalo & al. 2010, Ravera & Brunialti 2013), Pugl (Nimis & Tretiach 1999), Bas (Nimis & Tretiach 1999), Cal (Puntillo 1996, Incerti & Nimis 2006), Si (Schicchi & al. 1997, Falco Scampatelli 2005, Campisi & al. 2020).
Description: Thallus crustose, usually smooth, continuous to finely rimose in central parts, white to very pale grey, sometimes delimited by a dark prothallus. Apothecia lecanorine, 1-3 mm across, round or irregular in outline, slightly constricted at base, with an ochraceous brown to medium brown, at first slightly pruinose, flat to finally convex disc, and a thick, swollen, often flexuose, pruinose thalline margin. Thalline exciple without a differentiated cortex, the medullary hyphae reaching the margin, with abundant, small, yellowish crystals insoluble in K; epithecium orange to orange-brown, the pigment not dissolving in K, with coarse, yellowish crystals dissolving in K; hymenium colourless, 75-110 µm high; paraphyses slightly thickened at apex; hypothecium colourless, not inspersed. Asci 8-spored, clavate, very thin-walled, with a K/I+ blue, tall tholus penetrated by a faintly amyloid apical cushion, the wall K/I-, surrounded by a blue outer layer, Lecanora-type. Ascospores 1-celled, hyaline, ellipsoid, 11-18 x 5-8 µm. Photobiont chlorococcoid. Spot tests: thallus K+ yellow, C-, KC-, P+ yellow to orange-red (reaction more evident in the thalline margin of apothecia), UV+ pale orange; apothecial disc P-. Chemistry: atranorin (major), chloratranorin, psoromic acid, 2'-0-demethylpsoromic acid, zeorin.
Note: a cool-temperate species found on smooth subacid bark, with optimum in humid beech forests; present throughout the country, but most common in the beech forests of central and southern Italy.
Growth form: Crustose
Substrata: bark
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: very common
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis CC BY-SA 4.0
TSB 5254



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20157)
2001/12/08



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (20171)
2008.03.06



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5254)
2008.03.06



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08



Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08



Domenico Puntillo; Owner: Domenico Puntillo
Italy



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Passo del Pura, Ampezzo Carnico, 1400 m
09.09.2016



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Passo del Pura, Ampezzo Carnico, 1400 m
09.09.2016



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



Felix Schumm – CC BY-SA 4.0
[1006], Germany, Baden-Württemberg, Schwäbische Alb, Kreis Heidenheim, Reishalde beim Dudelhof südlich von Steinheim, ca. 630 m, Buchen im Laubwald, TK 7326/1. Leg. et det. Schumm.



P.L.Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5254)
2008.03.06



P.L. Nimis CC BY-SA 04
TSB 24363