Acarospora fuscata (Schrad.) Arnold

Flora, 53: 469, 1870. Basionym: Lichen fuscatus Schrad. - Spicil. Fl. Germ., 1: 83, 1794 nom. cons. prop.
Synonyms: Acarospora fuscata var. minutissima Bagl.; Acarospora monacensis H. Magn.?; Acarospora nigrocastanea Hue; Acarospora photina A. Massal.; Acarospora rufoalutacea Harm. ex H. Magn.?; Acarospora smaragdula sensu A. Massal. et auct. ital. p.p.; Lecanora cervina var. squamulosa (Schrad.) Willey
Distribution: N - VG, Frl (Tretiach & Hafellner 2000), Ven (Lazzarin 2000b), TAA (Caniglia & al. 2002, Nascimbene 2003, 2008b, Thor & Nascimbene 2007), Lomb (Brackel 2010, 2013), Piem (Morisi & Sereno 1995, Isocrono & Falletti 1999, Isocrono & al. 2003, 2004, 2006, Favero-Longo & al. 2006b, Isocrono & Piervittori 2008, Giordani & al. 2014), VA (Piervittori & Isocrono 1999, Valcuvia 2000, Matteucci & al. 2015c), Emil (Dalle Vedove & al. 2002, Tretiach & al. 2008, Fariselli & al. 2020), Lig (Watson 2014, Ravera & al. 2022). C - Tosc (Tretiach & al. 2008, Brackel 2015), Marc, Laz, Sar (Monte 1993, Nöske 2000, Rizzi & al. 2011, Giordani & al. 2013, Brackel & Berger 2019). S - Camp (Ricciardi & al. 2000), Pugl, Bas, Cal (Puntillo 1996), Si (Ottonello & Romano 1997, Poli & al. 1997, Grillo 1998, Grillo & Caniglia 2004, Brackel 2008b, Ottonello & al. 2011).
Description: Thallus crustose, episubstratic, areolate, often forming extensive, up to 10 cm wide patches. Areoles 0.2-2(-3) mm wide, up to 1 mm thick, mostly contiguous, irregular in outline, usually flat and uneven, epruinose, pale to reddish brown, often with black margins and undersides, epruinose. Cortex 25-35(-50) µm thick, yellow-brown in uppermost part, otherwise colourless, paraplectenchymatous, with perpendicularly arranged cells; algal layer continuous, 50-100 μm thick; medulla white, 0.1-0.8 mm thick, of intricate, thin-walled hyphae. Apothecia 1-5(-10) per areole, 0.1-0.4(-1) mm acrosss, at first punctiform, later expanded, angular to irregular in outline, with a concave to flat, usually angular or irregular, immersed, reddish brown to blackish disc, and an inapparent thalline margin. Proper exciple distinct, 10-20(-30) μm wide, not expanding around the disc; epithecium brownish, 10-15 μm high; hymenium colourless, (70-)100-120(-140) μm high, hemiamyloid, K/I+ red or blue turning red; paraphyses coherent, mostly simple, (1-)1.5-2 μm thick at mid-level, the apical cells barely expanded or 2-4 μm wide; hypothecium colourless, 30-60 μm high, euamyloid, K/I+ blue. Asci 100-200-spored, clavate, the apical dome K/I-. Ascospores 1-celled, hyaline, narrowly ellipsoid to subcylindrical, 4-6(-7) x 1-2 μm. Pycnidia immersed. Conidia 1-2 x 0.5-1 μm. Photobiont chlorococcoid. Spot tests: cortex K-, C+ red, KC+ red, P-, UV-; medulla C-, KC-. Chemistry: cortex with gyrophoric acid.
Note: a holarctic species of acid siliceous rocks wetted by rain, sometimes growing on other lichens, with a wide altitudinal range. Several records, especially those from southern Italy, need confirmation, but the species is certainly widespread throughout the country.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rather rare
Subalpine belt: rather rare
Oromediterranean belt: rare
Montane belt: rare
Submediterranean belt: very rare
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35221)
2001/12/12



Felix Schumm – CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.


Andres Saag; Owner: University of Tartu



Domenico Puntillo; Owner: Domenico Puntillo
Italy, Herb. Puntillo 4615



Juri Nascimbene - CC BY-SA 4.0
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern



Juri Nascimbene - CC BY-SA 4.0
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern



Juri Nascimbene - CC BY-SA 4.0
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern



Juri Nascimbene - CC BY-SA 4.0
Italy, Trentino-Alto Adige, Bolzano/Bozen, Sciliar/Schlern



Felix Schumm – CC BY-SA 4.0
[2636], Germany, Baden-Württemberg, Zollernalbkreis, an Kalkfelsen beim Lochenstein nahe Balingen, ca. 900 m, leg. et det. Schumm 26.06.1978.



Felix Schumm – CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (33464)
2001/12/12



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Felix Schumm CC BY-SA 4.0
[16527], Germany, Baden-Württemberg, Schwarzwald-Baar-Kreis, an der Straße von Hornberg nach Schramberg, Abbiegung Käppelehof. 48,22853° N, 8,31805° E, 670 m. An Granitfelsen. Leg. et det. Schumm 07.07.2010.



Pier Luigi Nimis CC BY-SA 4.0
TSB 41608



Pier Luigi Nimis CC BY-SA 4.0
TSB 41696



Pier Luigi Nimis CC BY-SA 4.0
TSB 41606



Pier Luigi Nimis - CC BY-SA 4.0
TSB 38642



Pier Luigi Nimis - CC BY-SA 4.0
TSB 38643



Pier Luigi Nimis - CC BY-SA 4.0
TSB 38642



Pier Luigi Nimis - CC BY-SA 4.0
TSB 38642