Lobarina scrobiculata (Scop.) Cromb.

Nyl. ex Cromb., Monogr. Lich. Brit. 1: 270, 1894. Basionym: Lichen scrobiculatus Scop. - Fl. Carn., ed. 2, 2: 384, 1772.
Synonyms: Lobaria scrobiculata (Scop.) DC.; Lobaria verrucosa (Huds.) Hoffm.; Lobaria verrucosa f. esorediosa Gyeln.; Lobarina scrobiculata (Scop.) Nyl.; Parmelia scrobiculata (Scop.) Ach.; Sticta scrobiculata (Scop.) Ach.
Distribution: N - Frl (Tretiach 1993, 1996), Ven (Lazzarin 1997, Nascimbene 2003b, 2011, Nascimbene & al. 2005b, 2006c, 2007, 2009c, 2010b), TAA (Nascimbene & Caniglia 2000b, Nascimbene & al. 2007b, Nimis & al. 2015), Lomb (Dalle Vedove & al. 2004), Piem (Isocrono & al. 2004, Morisi 2005), VA (Piervittori & Isocrono 1999, Matteucci & al. 2013), Emil (Tretiach & al. 2008, Fariselli & al. 2020), Lig (Brunialti & al. 1999, Giordani & Brunialti 2000, Watson 2014). C - Tosc (Tretiach & Nimis 1994, Benesperi & al. 2007, Brunialti & Frati 2010, Benesperi 2011, Brackel 2015), Marc (Nimis & Tretiach 1999), Umb (Ravera 1998, Ravera & al. 2006), Laz (Ravera 2002b), Abr (Nimis & Tretiach 1999), Mol (Caporale & al. 2008), Sar (Tretiach 1993, Zedda 1995, 2002, 2002b, Rizzi & al. 2011). S - Camp (Ricciardi & al. 2000, Tretiach 1993, Nascimbene & al. 2010b, Catalano & al. 2010, 2016, Ravera & Brunialti 2013), Pugl, Bas (Fascetti & al. 2005, Potenza 2006, Puntillo & al. 2009, Potenza & Fascetti 2010, 2012, Ravera & al. 2021c), Cal (Tretiach 1993, Puntillo & Vezda 1994, Puntillo 1995, 1996, Incerti & Nimis 2006), Si (Merlo 2004).
Description: Thallus foliose, broad-lobed, heteromerous, loosely attached, 10-16 cm wide. Lobes rounded at apex, contiguous to imbricate, 10-30 mm wide, with entire to slightly crenulate margins, blue-grey to yellowish grey when dry, dark blue-grey to blue-green when wet, undulate, coarsely reticulate-areolate but without defined ridges, sorediate. Soralia blue-grey, mainly laminal, punctiform at first, but eroding with age and becoming circular; soredia granular. Lower surface with glabrous, whitish patches separated by a pale brown, silky, entangled tomentum, erhizinate or with very sparse, simple rhizines. Upper and lower cortex paraplectenchymatous, the lower cortex poorly developed; medulla white. Apothecia very rare, 1-2 mm across, with a concave, red-brown disc and a verruculose, inflexed, often sorediate thalline margin. Asci 8-spored, fissitunicate, the thickened apex with a K/I+ blue ring, Peltigera-type. Ascospores 1-3-septate, hyaline, elongate-fusiform, 17-23 x 4-7 µm. Photobiont cyanobacterial (Nostoc in short chains). Spot tests: cortex K-, C-, KC-, P-; medulla K- or K+ yellow-orange, C-, KC- or KC+ pink-orange, P- or P+ orange, UV+ violet. Chemistry: chemotype 1 has usnic and stictic acids plus scrobiculin and sometimes traces of norstictic acid; chemotype 2 lacks stictic acid.
Note: a mild-temperate, suboceanic species found on old deciduous trees and on mossy rocks in humid open forests; formerly more frequent, presently extinct in several parts of the country (e.g. in the whole of the Po-plain), and declining elsewhere, with optimum in old Castanea-stands. A distribution map in Italy was published by Nascimbene & al. (2016). It is included in the Italian red list of epiphytic lichens as “Near-threatened” (Nascimbene & al. 2013c).
Growth form: Foliose, broad lobed
Substrata: bark
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis CC BY-SA 04
TSB 5109



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5109)
2001/11/26
detail of soredia



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5109)
2001/11/26
detail of lower surface with tomentum


Triin Aimla; Owner: University of Tartu
originally as Lobaria scrobiculata


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php



Juri Nascimbene - CC BY-SA 4.0
Italy, Veneto, Belluno, Val Visdende, Cadore
2007



Juri Nascimbene – CC BY-SA 4.0; Owner: Department of ùLife Sciences, University of Trieste
Vinca, Prov. Massa Carrara, Tuscany, Italy



Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3



Felix Schumm – CC BY-SA 4.0
[19194], Norway, Eidford, 60.469° N, 7.066° E, 40 m, growing on shady deciduous tree. Leg. F. Schumm 21.06.2014.



Felix Schumm – CC BY-SA 4.0
[19194], Norway, Eidford, 60.469° N, 7.066° E, 40 m, growing on shady deciduous tree. Leg. F. Schumm 21.06.2014.



Felix Schumm – CC BY-SA 4.0
[19194], Norway, Eidford, 60.469° N, 7.066° E, 40 m, growing on shady deciduous tree. Leg. F. Schumm 21.06.2014.



Felix Schumm – CC BY-SA 4.0
[19194], Norway, Eidford, 60.469° N, 7.066° E, 40 m, growing on shady deciduous tree. Leg. F. Schumm 21.06.2014.



Felix Schumm – CC BY-SA 4.0
[19194], Norway, Eidford, 60.469° N, 7.066° E, 40 m, growing on shady deciduous tree. Leg. F. Schumm 21.06.2014.



Felix Schumm – CC BY-SA 4.0
[19194], Norway, Eidford, 60.469° N, 7.066° E, 40 m, growing on shady deciduous tree. Leg. F. Schumm 21.06.2014.



Felix Schumm – CC BY-SA 4.0
[19194], Norway, Eidford, 60.469° N, 7.066° E, 40 m, growing on shady deciduous tree. Leg. F. Schumm 21.06.2014.



Felix Schumm – CC BY-SA 4.0
[19194], Norway, Eidford, 60.469° N, 7.066° E, 40 m, growing on shady deciduous tree. Leg. F. Schumm 21.06.2014.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.



Felix Schumm – CC BY-SA 4.0
[19071], Australia, New South Wales, Great Dividing Range, Tallaganda State Forest, 10 km E of Captains Flat, 35°38' S, 149°31' E, 1240 m, on base of Eucalyptus viminalis in wet sclerophyll forest. Leg. J.A. Elix (19028) & D. Verdon, 22.03.1985. LICH. AUSTRALASICI EXS. 133.


Nere Sorrentino
Spain, Galicia, Pontevedra, on Quercus


Nere Sorrentino
Spain, Galicia, Pontevedra, on Quercus


Nere Sorrentino
Spain, Galicia, Pontevedra, on Quercus


Nere Sorrentino
Spain, Galicia, Pontevedra, on Quercus


Nere Sorrentino
Spain, Galicia, Pontevedra, on Quercus


Helena Rodríguez
Spain, Galicia


Helena Rodríguez
Spain, Galicia


Helena Rodríguez
Spain, Galicia


Helena Rodríguez
Spain, Galicia


Helena Rodríguez
Spain, Galicia


Helena Rodríguez
Spain, Galicia



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5109)
2001/11/26


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/


Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum - http://lichens-of-noricum.uni-graz.at/