Lobothallia radiosa (Hoffm.) Hafellner

Acta Bot. Malacitana, 16: 138, 1991. Basionym: Lichen radiosus Hoffm. - Enum. Lich. Eur., tab. 4, 1874.
Synonyms: Aspicilia radiosa (Hoffm.) Poelt & Leuckert; Aspicilia subcircinata (Nyl.) Coppins; Lecanora circinata (Pers.) Ach.; Lecanora radiosa (Hoffm.) Schaer.; Lecanora subcandicans (Müll. Arg.) Stizenb.; Lecanora subcircinata Nyl.; Placodium circinatum (Pers.) Gray; Placodium radiosum (Hoffm.) Ach.; Placodium subcircinatum (Nyl.) Arnold; Psoroma circinatum (Pers.) Rabenh.; Squamaria subcircinata (Nyl.) H. Olivier
Distribution: N - VG (Nimis & Tretiach 1995, Tretiach & Pecchiari 1995, Navarro-Rosinés & Hafellner 1996, Geletti 1997, Castello 2002, Martellos & Castello 2004, Tretiach & al. 2007b), Frl (Nimis & Salvadori 1998, Nascimbene & al. 2009b), Ven (Nascimbene & Caniglia 1997, Caniglia & al. 1999, Nascimbene 2005c, Nascimbene & Marini 2007), TAA (De Benetti & Caniglia 1993, Navarro-Rosinés & Hafellner 1996, Nascimbene 2003, Spitale & Nascimbene 2012), Lomb, Piem (Isocrono & al. 2004), VA (Piervittori & Isocrono 1999, Piervittori & al. 2001, 2004, Matteucci & al. 2008c, 2015c), Emil (Navarro-Rosinés & Hafellner 1996, Nimis & al. 1996, Benesperi 2009, Fariselli & al. 2020), Lig (Navarro-Rosinés & Hafellner 1996, Valcuvia & al. 2000, Giordani & al. 2016). C - Tosc (Adamo & al. 1993, Tretiach & Nimis 1994, Adamo 1997, Tretiach & al. 2008, Benesperi 2011, Brackel 2015), Marc (Nimis & Tretiach 1999), Umb (Nimis & Tretiach 1999, Panfili 2000b, 2007, Ravera & al. 2006, Genovesi 2011), Laz (Bartoli 1997b, Nimis & Tretiach 2004, Roccardi 2011, Brackel 2015), Abr (Nimis & Tretiach 1999, Caporale & al. 2016, Gheza & al. 2021), Mol (Nimis & Tretiach, 1999, 2004, Caporale & al. 2008, Genovesi & Ravera 2014, Brackel 2020), Sar (Rizzi & al. 2011, Giordani & al. 2013, Cossu & al. 2015, Brackel & Berger 2019). S - Camp (Garofalo & al. 1999, 2010, Aprile & al. 2002, 2003, 2003b, Nimis & Tretiach 2004, Catalano & al. 2016), Pugl (Nimis & Tretiach 1999, Gianfreda & Matino 2020), Bas (Nimis & Tretiach 1999, Brackel 2011), Cal (Puntillo 1996), Si (Nimis & al. 1994, 1996b, Ottonello & Salone 1994, Ottonello & al. 1994, 2011, Ottonello 1996, Navarro-Rosinés & Hafellner 1996, Ottonello & Romano 1997, Grillo 1998, Grillo & al. 2001, Grillo & Caniglia 2004, Merlo 2004b, Caniglia & Grillo 2005, 2006, Brackel 2008b, Gianguzzi & al. 2009, Liistro & Cataldo 2011, Cataldo & Cannavò 2014).
Description: Thallus crustose-placodioid, episubstratic, forming up to 6(-8) cm wide, orbicular rosettes, tightly adnate throughout, rimose to areolate in central parts, distinctly lobed at margins, pale to dark grey. Lobes contiguous, 3-7 x 0.4-1(-1.5) mm, c. 0.25 mm thick, sometimes divided and transversely broken by partial cracks, flat to slightly convex, sometimes weakly pruinose at tips. Cortex cellular, of anticlinally arranged hyphae, with abundant, fine, dark brown granules and hyaline, coarsely granular crystals, both insoluble in K; medulla white, with crystals, I-. Apothecia frequent, lecanorine-aspicilioid, mostly crowded in the center of thallus, long remaining immersed, finally adnate, 0.4-1.7 mm across, often compound or confluent and irregular in outline, with a red-brown to black disc and a thin, scarcely raised thalline margin. Epithecium brown to (rarely) olive-brown; hymenium colourless, K/I+ blue; paraphyses usually simple, tightly compacted, with globose, 2-3 µm wide apical cells; hypothecium colourless. Asci 8-spored, oblong-clavate, the thin outer coat K/I+ blue, the wall and apical dome K/I-. Ascospores 1-celled, hyaline, subglobose to broadly ellipsoid, (8-)11-13(-15) x 6-9(-11) µm. Pycnidia appearing as black dots, immersed, the wall colourless in section except in the periostiolar region. Conidia bacilliform, 4-6 x c. 1 µm. Photobiont chlorococcoid. Spot tests: medulla K- or K+ yellow turning red (needle-like crystals), C-, KC-, P- or P+ yellow-orange. Chemistry: medulla sometimes with norstictic acid, more rarely with stictic acid.
Note: a widespread holarctic lichen with a very wide altitudinal and latitudinal range, and with correspondingly broad ecological requirements, found on a wide variety of substrata, including basic siliceous rocks, limestone, dolomite, more rarely brick, roofing tiles and mortar; common throughout the country. The species, in its present circumscription, is chemically variable: the forms with norstictic acid, corresponding to “Aspicilia subcircinata”, may represent just a chemotype (see e.g. Roux & coll. 2014). See also note on L. parasitica.
Growth form: Crustose placodiomorph
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rather rare
Subalpine belt: rather common
Oromediterranean belt: common
Montane belt: very common
Submediterranean belt: extremely common
Padanian area: rather rare
Humid submediterranean belt: extremely common
Humid mediterranean belt: common
Dry mediterranean belt: rather common

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis CC BY-SA 4.0
TSB 22863



A. Moro; Owner: Department of Life Sciences, University of Trieste
Sicilia, TP, Isola di Marettimo
2008.04.03



A. Moro; Owner: Department of Life Sciences, University of Trieste
Sicilia, TP, Isola di Marettimo
2008.04.03



A. Moro; Owner: Department of Life Sciences, University of Trieste
Sicilia, TP, Isola di Marettimo
2008.04.03


Paolo Fontanesi, Reggio Emilia; Owner: Paolo Fontanesi, Reggio Emilia


Paolo Fontanesi, Reggio Emilia; Owner: Paolo Fontanesi, Reggio Emilia



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino (Rupingrande)
07.07.2016



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017



Pier Luigi Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Val Rosandra, San Lorenzo near Belvedere
11/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017



Andrea Moro; Owner: Department of Life Sciences, University of Triedte
Italy, Friuli Venezia Giulia, Trieste, Trieste Karst, Rocca di Monrupino
20/02/2017



Felix Schumm – CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm – CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm – CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm – CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm – CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm – CC BY-SA 4.0
[12290], Griechenland, N-Samos, bei Manolates südlich von Agios Konstandinos, 37°47.185'N, 26°49.533'E, 341 m; halbschattige bis besonnte Terrassenmauern. Leg. Schumm 02.05.2006



Felix Schumm – CC BY-SA 4.0
[12290], Griechenland, N-Samos, bei Manolates südlich von Agios Konstandinos, 37°47.185'N, 26°49.533'E, 341 m; halbschattige bis besonnte Terrassenmauern. Leg. Schumm 02.05.2006



Felix Schumm – CC BY-SA 4.0
[12290], Griechenland, N-Samos, bei Manolates südlich von Agios Konstandinos, 37°47.185'N, 26°49.533'E, 341 m; halbschattige bis besonnte Terrassenmauern. Leg. Schumm 02.05.2006



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm – CC BY-SA 4.0
[12290], Griechenland, N-Samos, bei Manolates südlich von Agios Konstandinos, 37°47.185'N, 26°49.533'E, 341 m; halbschattige bis besonnte Terrassenmauern. Leg. Schumm 02.05.2006



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm – CC BY-SA 4.0
[12290], Griechenland, N-Samos, bei Manolates südlich von Agios Konstandinos, 37°47.185'N, 26°49.533'E, 341 m; halbschattige bis besonnte Terrassenmauern. Leg. Schumm 02.05.2006



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm – CC BY-SA 4.0
[12290], Griechenland, N-Samos, bei Manolates südlich von Agios Konstandinos, 37°47.185'N, 26°49.533'E, 341 m; halbschattige bis besonnte Terrassenmauern. Leg. Schumm 02.05.2006



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010



Felix Schumm CC BY-SA 4.0
[16504], Germany, Baden-Württemberg, Kreis Göppingen, Gosbach- Bad Ditzenbach, Tierstein, 48,57461° N, 9,68471° E, 737 m, auf Kalkfelsen am Albrand. Leg. et det. Schumm 20.06.2010


Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php . Courtesy: Anita Stridvall


Jennifer Fiorentino
Malta



P.L. Nimis CC BY-SA 4.0
TSB 22863