Lepra excludens (Nyl.) Hafellner

in Hafellner & Türk, Stapfia, 104: 171, 2016. Basionym: Pertusaria excludens Nyl. - Flora, 68: 296, 1885.
Synonyms: Marfloraea excludens (Nyl.) S. Y. Kondr., L. Lőkös & J.-S. Hur; Pertusaria excludens Nyl.; Pertusaria inopinata Erichsen?
Distribution: N - Frl (Tretiach & Hafellner 2000), VA (Isocrono & al. 2008, Matteucci & al. 2015d), Lig. C - Tosc, Sar (Nöske 2000, Craighero 2010, Rizzi & al. 2011). S - Si (Craighero 2010).
Description: Thallus crustose, episubstratic, rather thick, markedly cracked and areolate, whitish grey, matt, sorediate, often delimited by a thin, pale prothallus. Areoles flat to slightly convex, irregular in outline, more or less angulose, 0.2-1(-2) mm wide, finely granulose or papillate. Soralia abundant, 0.6-1(-1.5) mm across, convex to excavate, slightly paler than the areoles and often with a yellowish tinge, discrete to finally confluent, bursting from the thalline warts and covering almost the whole thallus surface; soredia coarsely granular. Apothecia extremely rare, often aborting, hidden beneath the soredia. Ascospores unknown. Photobiont chlorococcoid. Spot tests: soralia K+ yellow turning blood red, C-, KC+ yellow-red, P+ yellow-orange, UV+ pale grey or grey-yellow. Chemistry: norstictic acid (major), salazinic acid (minor or traces), plus a UV+ yellow xanthone.
Note: on steeply inclined surfaces of sheltered siliceous rocks, probably more widespread also in the Alps.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: very rare
Dry mediterranean belt: absent

pH of the substrata:


Solar irradiation:








Altitudinal distribution:


Predictive model
Herbarium samples

P.L. Nimis CC BY-SA 4.0
TSB 17036

Nere Sorrentino
Spain, Canary Islands, Tenerife

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (17036)