in Hafellner & Türk, Stapfia, 104: 172, 2016. Basionym: Variolaria multipuncta Turner - Trans. Linn. Soc. London, 9: 137, 1808.
Synonyms: Pertusaria globulifera var. sorediata Mudd; Pertusaria leptospora Nitschke; Pertusaria multipuncta (Turner) Nyl.; Pertusaria sorediata C. Knight
Distribution: N - Frl, Ven (Nascimbene & Caniglia 2000), TAA (Nascimbene & al. 2007b), Lomb, Piem (Jatta 1909-1911), VA (Piervittori & Isocrono 1999), Emil (Fariselli & al. 2020). C - Tosc (Jatta 1909-1911), Abr (Di Santo & Ravera 2012, Corona & al. 2016), Mol (Ravera & Genovesi 2010, Ravera & al. 2010), Sar (Rizzi & al. 2011). S - Cal (Puntillo 1996, Incerti & Nimis 2006).
Description: Thallus crustose, episubstratic, ash grey, 1-3 mm thick, sometimes thinner, continuous to rimose, coarsely warted, usually well delimited and with a pale prothallus. Fruiting warts 0.5-1.5(-2) mm across, flattened, covered in white granular soredia. Apothecia immersed in the fruting warts (and hence visible only when these are sectioned), 1(-3) per wart, with a pale yellowish brown, pruinose, c. 0.5(-1) mm wide disc. Asci 1-spored, cylindrical, with thick, amyloid walls, Pertusaria-type. Ascospores 1-celled, elongate-ellipsoid to cylindrical, 90-200 × 30-70 μm, the wall to 10 μm thick, uniform. Photobiont chlorococcoid. Spot tests: thallus and soralia K+ yellow-brown, C-, KC+ yellow, P+ orange-red, UV- or faintly glaucous. Chemistry: physodalic acid (major), protocetraric acid (traces).
Note: a mainly temperate species found on smooth bark of deciduous trees (especially Carpinus and Fagus) in open, humid deciduous woodlands.
Growth form: Crustose
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)