Arthopyrenia analepta (Ach.) A. Massal.

Ric. Auton. Lich. Crost.: 165, 1852. Basionym: Lichen analeptus Ach. - Lichenogr. Suec. Prodr.: 15, 1799.
Synonyms: Arthopyrenia analepta f. mespili A. Massal.; Arthopyrenia analeptella (Nyl.) Arnold; Arthopyrenia arnoldii Zahlbr.?; Arthopyrenia fallax (Nyl.) Arnold; Arthopyrenia lapponina Anzi; Didymella fallax (Nyl.) Vain.; Leiophloea fallax (Nyl.) Riedl; Pseudosagedia fallax (Nyl.) Oxner; Sphaerella lapponum De Not.; Verrucaria epidermidis var. fallax Nyl.
Distribution: N - VG, Frl (TSB 5457), Ven (Lazzarin 2000b), TAA (Nascimbene & al. 2007b), Lomb (Brusoni & al. 1997, Brusoni & Valcuvia 2000, Anderi & al. 2005), Piem (Isocrono & al. 2004), Emil (Valcuvia & Grieco 1995, Fariselli & al. 2020), Lig. C - Marc, Laz, Mol (Caporale & al. 2018, Caporale & Ravera 2020). S - Camp (Nimis & Tretiach 2004, Garofalo & al. 2010), Bas (Potenza & al. 2014).
Description: Thallus crustose, inconspicuous, not discolouring the bark, not lichenized. Perithecia black, usually ellipsoid, with a flat border formed by the spreading involucrellum (0.3-)0.4-0.6 x 0.2-0.5 mm. Involucrellum dimidiate, brown in water, K+ greenish, of compacted hyphae and bark cells; exciple thin, usually colourless, surrounding the centrum; pseudoparaphyses slender, distantly branched, 1.5-2 μm thick, the cells 8-15(-18) μm long; downwardly projecting periphysoids present near the ostiole. Asci 8-spored, subcylindrical, fissitunicate, with two functional wall layers, 65-100 x 12-16(-20)μm. Ascospores 1-septate (a few old spores occasionally 3-septate), clavate with rounded apices, strongly constricted at septum, with often unequal cells, each cell often biguttulate but without a median constriction (except sometimes in the lower cell), (13-)15-23(-24) x 5.5-8(-9) μm; perispore distinct, c. 2 μm thick in K. Pycnidia black, not always present, 50-100 μm across. Conidia hyaline, thread-like, more or less straight, 7-11 x 0.8-1 μm. Photobiont absent. Spot tests: all negative. Chemistry: without lichen substances.
Note: a mainly temperate, perhaps holarctic early coloniser of smooth bark, found on twigs of deciduous trees, especially Carpinus and Corylus, but also on Quercus and Sorbus, facultatively, or most probably non-lichenised. For nomenclatural matters see Harris (1995).
Growth form: Fungus
Substrata: bark
Reproductive strategy: mainly sexual
Pioneer species

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: extremely rare
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: very rare
Padanian area: extremely rare
Humid submediterranean belt: rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:


Solar irradiation:








Altitudinal distribution:


Predictive model
Herbarium samples

Walter Obermayer CC BY-SA 4.0 - Source: Lichens of Noricum -

Sonia Ravera
Det. B. Coppins

Paolo Giordani

Deborah Isocrono

Photo uploaded by P. Cannon - CC BY-SA NC - Source:

Paolo Giordani

Paolo Giordani

Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.

G. Gibelli, in S. Garovaglio, 1865 Tentamen Dispositionis Methodicae Lichenum In Longobardia Nascentium. Mediolanum