Leptorhaphis atomaria (Ach.) Szatala

Magyar Bot. Lapok, 26: 31, 1928 ("1927"). Basionym: Lichen atomarius Ach. - Lichenogr. Suec. Prodr.: 16, 1799.
Synonyms: Arthopyrenia analepta var. atomaria (Ach.) Jatta; Arthopyrenia atomaria (Ach.) Müll. Arg.; Arthopyrenia persoonii f. atomaria (Ach.) A. Massal.; Arthopyrenia punctiformis var. atomaria (Ach.) Anzi; Didymella atomaria (Ach.) Szatala; Leiophloea punctiformis var. atomaria (Ach.) Trevis.; Leptorhaphis tremulae auct. ital. non Körb.; Microthelia adspersa Körb.; Sagedia salicis A. Massal.?; Verrucaria epidermidis var. atomaria (Ach.) Schaer.; Verrucaria populicola Nyl.?; Verrucaria punctiformis var. atomaria (Ach.) Ach.
Distribution: N - VG (TSB 2345), Ven (Lazzarin 2000b), TAA (Nascimbene & al. 2007b), Lomb (Brusoni & al. 1997, Brusoni & Valcuvia 2000), Piem (Isocrono & al. 2004), Lig (Giordani & al. 2009). C - Tosc. S - Cal (Puntillo & Puntillo 2004), Si (Ottonello & al. 2011).
Description: Thallus non- or doubtfully lichenized, crustose, mostly endosubstratic, appearing as a pale grey to white, continuous, poorly delimited patch. Perithecia black, matt, scattered, 0.1-0.25 µm across, flattened, with a central ostiole. Involucrellum greenish-brown, dimidiate, sometimes extending beyond the perithecium into a thin fringe, consisting of bark cells mixed with greenish-black, thick-walled, isodiametric cells with large lumina; exciple colourless to pale brown, of thin-walled, flattened cells; pseudoparaphyses sparse, c. 2 µm thick; hymenial gel I+ blue. Asci (6-)8-spored, clavate, bi- and fissitunicate, with a rounded to truncate apex. Ascospores 1-3-septate, hyaline, falcate or sigmoid, with more or less rounded ends, (15-)25-32(-35) x 2-3.5 µm, without epispore. Pycnidia black, subglobose, scattered, at first immersed, then projecting. Conidia 1-celled, narrowly fusiform, curved, (11.5-)16.5-18 x 1.5-2.5 µm. Photobiont absent or trentepohlioid. Spot tests: K-, C- KC-, P-. Chemistry: without lichen substances.
Note: a doubtflully lichenised species, found on base-rich bark of deciduous trees, mainly Populus tremula, Salix and Fraxinus. Probably overlooked, and more widespread.
Growth form: Fungus
Substrata: bark
Reproductive strategy: mainly sexual
Pioneer species

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: rather rare
Padanian area: absent
Humid submediterranean belt: rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

12345

Solar irradiation:

12345

Aridity:

12345

Eutrophication:

12345

Poleotolerance:

0123

Altitudinal distribution:

123456


Predictive model
Herbarium samples


Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m.
Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (9183)
2001/11/26



Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m. Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011



Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m. Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011



Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m. Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011



Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m. Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011



Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m. Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011



Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m. Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011



Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m. Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011


Photo by Maria Zardini, Museo di Storia Naturale di Venezia CC BY-SA 4.0
Trevisan Lichenotheca Veneta 252



Felix Schumm – CC BY-SA 4.0
[13695], Malta, Xemxija, Pinus halepensis Wäldchen, 35°56.760'N, 14°22.193'E, 53 m. Leg. Schumm, Stapper, Lüth und Frahm 12.03.2008, det. Aptroot 2011