Description: Thallus squamulose, the squamules pale to dark brown, scattered or contiguous, 1.5-6(-8) mm wide, 0.15-0.4 mm thick, rounded to deeply lobate, flat and fully adnate or concave with upturned, often undulate, slightly thickned and black-rimmed margins; lower surface brown-black, attached by conspicuous, up to 10 mm long, black, branched rhizines, without rhizohyphae. Upper cortex 30-50 µm thick, of angular, isodiametric or anticlinally slightly elongated cells, overlain by a 10-30 µm thick epinecral layer; medulla white, of interwoven hyphae; algal layer c. 50-80 µm thick, the algal cells arranged in vertical columns; lower cortex paraplectenchymatous, brown-black, 25-40(-50) µm thick. Perithecia black, broadly pyriform, immersed, up to 0.4 mm across (in section). Exciple black, often paler near the ostiole; paraphyses absent, substituted by 40-70 µm long periphyses; hymenium colourless, I+ brown-red; hymenial algae globose, 3-4 μm wide or ellipsoid and up to 5 x 3 μm. Asci 1-spored, clavate to cylindrical-clavate, bi- and fissitunicate, thin-walled, the wall non-amyloid. Ascospores muriform, yellowish brown to brown, elongate-ellipsoid or narrowly clavate, (47-)65-90(-110) x 23-36 µm. Pycnidia immersed, up to 0.2 mm across. Conidia bacilliform, 3-4.5 x c. 0.8 µm. Photobiont chlorococcoid (probably Diplosphaera, present in both thallus and hymenium). Spot tests: thallus and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a widespread terricolous species also known from the Southern Hemisphere, growing on clay soils in opening of Mediterranean garrigue vegetation. Probably more widespread in Mediterranean Italy.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris