Description: Thallus crustose, episubstratic, grey-green to dark green, warted-areolate. Areoles usually convex, incised, almost subsquamulose, 0.1-0.25 mm wide. Soralia yellowish green to pale green, paler than thallus, at first well-delimited and rounded, 0.25-0.7 mm across, but soon confluent and covering the central parts of thallus; soredia 15-30(-40) µm in diam., often forming larger consoredia (50-75 µm in diam). Cortex poorly developed, c. 10 µm thick. Apothecia rare, biatorine, rounded to irregular in outline, sessile with a constricted base, 0.4-1 mm across, with a pale pink to red-brown, epruinose, convex or tuberculate disc, and a thin, slightly paler, soon excluded proper margin. Proper exciple of radiating, hyaline hyphae, 50-95 μm wide; epithecium poorly differentiated from the hymenium; hymenium colourless, 40-55 µm high, C-; paraphyses simple, 1.5-2 µm thick at base, apically slightly swollen and 2-3 µm thick; hypothecium colourless or pale yellowish brown, up to 300 µm thick. Asci 8-spored, clavate, with a K/I+ blue apical dome penetrated by a narrow, K/I– apical cushion surrounded by a narrow, deeply K/I+ blue zone, the wall K/I- but surrounded by an I+ red-brown, K/I+ blue outer layer, the ocular chamber relatively small, Biatora-type. Ascospores 0(-1)-septate, hyaline, narrowly ellipsoid, (10-)12-16(-20) x (3-)4-6(-7) µm. Photobiont chlorococcoid. Spot tests: thallus and soralia K-, C+ red, KC+ red, P-. Chemistry: gyrophoric acid (major), lecanoric acid (accessory).
Note: on epiphytic bryophytes in humid forests, mostly in upland areas.
Growth form: Crustose
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)