Calogaya bryochrysion (Poelt) Vondrák

in Vondrák & al., Lichenologist, 48: 177, 2016. Basionym: Caloplaca bryochrysion Poelt - Feddes Rep., 58: 175, 1955.
Synonyms:
Distribution: N - Frl, Ven (Nimis 1994, Caniglia & al. 1999), TAA (Nascimbene 2003), Piem. C - Marc (Nimis & Tretiach 1999), Abr (Gheza & al. 2021).
Description: Thallus crustose, of scattered or contiguous, circular to irregular, weakly to strongly convex, 0.1-0.5(-0.8) mm wide, 0.1-0.2 mm thick areoles, yellow, yellowish orange to orange, forming small irregular patches. Soralia at first punctiform to small-crateriform and laminal, yellow to orange, later becoming convex and covering most of the thallus; soredia granular, 20-35 μm in diam. Apothecia usually absent, or sparse and scattered, 0.3-0.6(-1) mm across., sessile, slightly to strongly constricted at base, more or less zeorine with a concave to strongly convex, yellowish orange to orange, smooth, epruinose disc, and an inconspicuous, often excluded thalline margin; proper margin raised or level with disc, slightly paler, smooth. Epithecium yellow to yellowish orange, K+ red; hymenium colourless, 55-80 μm thick; paraphyses not gelatinized, simple to branched, the terminal cells swollen, up to 5 μm wide, often with oil-droplets; hypothecium colourless, with oil-droplets. Asci (6-)8-spored, clavate, functionally unitunicate, apically thickened with a broad internal beak, the inner part of apex and external cap I+ blue, Teloschistes-type. Ascospores 2-celled, polarilocular, hyaline, broadly ellipsoid, (7-)8-12(-18) x (4-)5.5-8(-9) μm, the equatorial thickening (“septum”) 2-3.5 μm. Photobiont chlorococcoid. Spot tests: thallus and apothecia K+ dark red, C-, KC-, P-. Chemistry: thallus and apothecia with emodin, fallacinal, parietin, parietinic acid, and teloschistin.
Note: on mosses, soil and plant debris over calcareous substrata, but also directly on calcareous rocks, in sheltered but light-rich situations, with optimum above treeline. Very much overlooked, or confused with other sorediate species in the Alps, it occurs also in Abruzzo (Majella Massif). For further details see Vondrák & al. (2016b).
Growth form: Crustose
Substrata: rocks, soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)

Commonnes-rarity: (info)

Alpine belt: rare
Subalpine belt: very rare
Oromediterranean belt: extremely rare
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Felix Schumm - CC BY-SA 4.0
[BR5030040676335], Austria, Tirol, Ötztal, Ober-Gurgel, Zirbelwald and Spitzkögele, in alpina heathland, 2000-2100 m. Leg. A. Aptroot (14072), 14-17.1984, det. A. Aptroot, 1986.



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (8703)
2001/12/18


Felix Schumm - CC BY-SA 4.0
[BR5030040676335], Austria, Tirol, Ötztal, Ober-Gurgel, Zirbelwald and Spitzkögele, in alpina heathland, 2000-2100 m. Leg. A. Aptroot (14072), 14-17.1984, det. A. Aptroot, 1986.


Felix Schumm - CC BY-SA 4.0
[BR5030040676335], Austria, Tirol, Ötztal, Ober-Gurgel, Zirbelwald and Spitzkögele, in alpina heathland, 2000-2100 m. Leg. A. Aptroot (14072), 14-17.1984, det. A. Aptroot, 1986.



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0068097], Spain, prov. Gerona, Pyrenean mountains, Nuria, 15 km E of Puigcerda, c 1 km NE of the Monastery, weakly calcareous rocks in alpine meadow on NW-facing slope, in fissure in steep rockface, 2300 m. Leg. H. Sipman (No.8156), 7.09.1976, det. N. Ernste, 1977



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0068097], Spain, prov. Gerona, Pyrenean mountains, Nuria, 15 km E of Puigcerda, c 1 km NE of the Monastery, weakly calcareous rocks in alpine meadow on NW-facing slope, in fissure in steep rockface, 2300 m. Leg. H. Sipman (No.8156), 7.09.1976, det. N. Ernste, 1977



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0068097], Spain, prov. Gerona, Pyrenean mountains, Nuria, 15 km E of Puigcerda, c 1 km NE of the Monastery, weakly calcareous rocks in alpine meadow on NW-facing slope, in fissure in steep rockface, 2300 m. Leg. H. Sipman (No.8156), 7.09.1976, det. N. Ernste, 1977



P.L. Nimis CC BY-SA 04
TSB 37151



P.L. Nimis CC BY-SA 04
TSB 37152