Kuettlingeria cretensis (Zahlbr.) I.V. Frolov & Vondrák

in Frolov & al., J. Syst. Evol. 59, 3: 468, 2020. Basionym: Blastenia cretensis Zahlbr. - Sitzungsber. K. Akad. Wiss. Wien, math.-naturw. Kl., 115: 519, 1906.
Synonyms: Blastenia ulbensis Zahlbr.; Caloplaca calcicola Zahlbr.; Caloplaca cretensis (Zahlbr.) Wunder
Distribution: C - Mol (Nimis & Tretiach 1999, Caporale & al. 2008). S - Camp (Nimis & Tretiach 2004, Garofalo & al. 2010), Si (Nimis & al. 1994, Grillo & al. 2007).
Description: Thallus crustose, endosubstratic and poorly evident, or thinly episubstratic and finely rimose-areolate, whitish, ochraceous, brownish or grey, rarely delimited by a black prothallus. Apothecia biatorine/zeorine, sessile and constricted at base, round, (0.1-)0.2-0.5(-0.6) mm across, with an orange, brownish orange to black, often orange-brown-pruinose, mostly flat to finally slightly convex disc, and an initially thick and smooth, finally often excluded, at first orange, then often blackening proper margin; on the same thallus the apothecia vary form completely orange to completely black, the blackening usually starting from the disc. Thalline exciple sometimes developed only at the base of the apothecia; proper exciple up to 100 µm thick, the outer part corticate, the inner part colourless; epithecium pale violet grey, with a granular, yellow-orange epipsamma reacting K+ purplish-red, the non-granular parts reacting K+ violet; hymenium colourless, 60-120 µm high, not inspersed with oil droplets; paraphyses coherent, 1.5-2.5 µm thick at mid-level, the apical cells 2-4(-6) µm wide; hypothecium colourless, not inspersed with oil droplets, 60-120 µm high. Asci 8-spored, clavate, Teloschistes-type. Ascospores 1-septate, polarilocular, hyaline, ellipsoid, 10-17 x 4-9.5 µm, the septum 2-3.5(-5) µm thick. Photobiont chlorococcoid. Spot tests: Thallus K- or K+ violet, C-, KC-, P-. Apothecia K+ red/violet. Chemistry: apothecia and sometimes thallus with the Sedifolia-grey pigment (blackened parts); apothecia with additional anthraquinones.
Note: an (apparently) eastern Mediterranean species also known from southern France (Roux & coll. 2014), found on compact limestones, mostly coastal in Italy; probably more widespread in the Mediterranean Region than the few records would suggest.
Growth form: Crustose
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Taxon bound to maritime-coastal situations

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: extremely rare
Dry mediterranean belt: extremely rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (18656)
2001/12/18



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (18656)
2001/11/19



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



Felix Schumm; Owner: Felix Schumm - CC BY-SA 4.0
[B 60 0135247], Greece, W Aegean, Nomos Evvias, Eparchia Chalkidos, Dimos Steni Dirfios, Central Evvia, Chiliadou beach at E coast, N of Stropones, limestone rocks and cliffs in phrygana near the coast 38°40.3' N, 23°55.2' E, 10 m. Leg. H. Sipman (54439) & Th. Raus, 3. October 2005, det. H. Sipman, 2006



P.L. Nimis CC BY-SA 04
TSB 36923



P.L. Nimis CC BY-SA 04
TSB 36923



P.L. Nimis CC BY-SA 04
TSB 36923



P.L. Nimis CC BY-SA 04
TSB 36923



P.L. Nimis CC BY-SA 04
TSB 36923



P.L. Nimis CC BY-SA 04
TSB 36923