Catillaria contristans (Nyl.) Zahlbr.

Cat. Lich. Univ., 4: 35, 1926. Basionym: Lecidea contristans Nyl. - Flora, 48: 354, 1865.
Synonyms: Biatora hypocyanea (Stirt.) Zahlbr.; Biatorina contristans (Nyl.) Arnold; Biatorina sphaeralis (Körb.) Jatta; Catillaria dufourii (Ach. ex Nyl.) Vain.; Catillaria sphaeralis Körb.; Lecidea dufourii Ach. ex Nyl.; Lecidea hypocyanea Stirt.; Lecidea sabuletorum f. simplicior Nyl.
Distribution: N - Lomb.
Description: Thallus crustose, episubstratic, rather thick, warty-granular, often cracked, whitish to dark grey or grey-brown, without a distinct prothallus, forming irregular, spreading, up to 2 cm wide patches. Areoles/granules 0.05-0.3(0.5) mm wide, convex to subglobose, contiguous. Apothecia biatorine, black, often somehow shiny, crowded and sometimes confluent, 0.2-0.6(-1) mm across, with a strongly convex, epruinose disc, and a thin, soon excluded proper margin which is sometimes surrounded by a white byssoid rim. Proper exciple colourless or greenish brown in outer part, of coherent hyphae; epithecium dark green to olive-green, K-. N+ red; hymenium colourless to greenish in upper part and pale brown in lower part, 35-60 µm high; paraphyses coherent, simple or with numerous short branches, 1.5-2.5(-3) µm thick at mid-level, the apical cells not markedly swollen; hypothecium appearing colourless to pale brown in thin sections, dark reddish brown to brown-black in thick sections. Asci 8-spored, clavate Biatora-type. Ascospores (0-)1-septate, constricted at septum, hyaline, ellipsoid to ovoid-oblong, 9-16.5 x 2.5-4.5 µm. Pycnidia black, immersed. Conidia bacilliform. Photobiont chlorococcoid, the cells 9-16 µm in diam. Spot tests: thallus K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: on dead bryophytes (Andreaea, Grimmia) and soil rich in humus over acid siliceous rocks in upland areas. According to Coppins (1983) and Kistenich & al. (2018), it does not belong to Catillaria.
Growth form: Crustose
Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: very rare
Subalpine belt: very rare
Oromediterranean belt: absent
Montane belt: very rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:


Solar irradiation:








Altitudinal distribution:


Predictive model

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb. Poelt (2-93)

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb. Poelt (2-93)

Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.