Leptogium cyanescens (Ach.) Körb.

Syst. Lich. Germ.: 420, 1855. Basionym: Collema tremelloides f. cyanescens Ach. - Syn. Meth. Lich.: 326, 1814.
Synonyms: Leptogium caesium (Ach.) Vain.
Distribution: N - VG (TSB 10787), Frl, Ven (Nascimbene & al. 2005b, 2006c, 2007, Nascimbene & Marini 2010), TAA (Nascimbene & al. 2006e, 2007b), Lomb, Piem (Isocrono & al. 2004, Matteucci & al. 2013), VA (Piervittori & Isocrono 1999, Piervittori & al. 2001), Emil (Fariselli & al. 2020), Lig (Brunialti & al. 1999). C - Tosc, Marc, Laz (Ravera 2001, Massari & Ravera 2002), Abr (Ravera 2002b, Corona & al. 2016), Mol (Paoli & al. 2015, Caporale & Ravera 2020), Sar (Zedda 2002). S - Camp (Ricciardi & al. 2000, Nimis & Tretiach 2004, Garofalo & al. 2010, Catalano & al. 2010, 2016), Pugl, Cal (Puntillo 1995, 1996), Si (Ottonello & Romano 1997, Grillo & al. 2002, Grillo & Caniglia 2004, Caniglia & al. 2005, Caniglia & Grillo 2006b, Ottonello & Puntillo 2009, Ottonello & al. 2011).
Description: Thallus foliose, subgelatinous when wet, 1-5(-10) cm in diam., adnate. Lobes irregular, elongate, plane, separate, 4-5(-10) mm wide, 35-100 µm thick, entire, dentate to irregularly cut and isidiate, with rotund apices. Upper surface blue-grey, smooth to somewhat wrinkled, with dense, laminal to marginal, usually cylindrical or flattened isidia. Upper and lower cortices consisting of a single layer of isodiametrical cells, internally with loosely interwoven chains of Nostoc and hyphae. Lower surface grey, smooth, with scattered tufts of white hairs but not densely tomentose. Apothecia rare, laminal, sessile to substipitate, 0.5-2 mm wide, with a brown disc and an often isidiate thalline margin. Proper exciple euparaplectenchymatous; epithecium pale brown; hymenium colourless; paraphyses simple, c. 1 µm wide, slightly inflated at apex; hypothecium colourless. Asci: 8-spored, cylindrical-clavate. Ascospores submuriform, hyaline, (15-)18-23(-25) x (6-)7-9(-10) µm. Photobiont cyanobacterial (Nostoc), in long chains. Spot tests: all negative. Chemistry: without lichen substances.
Note: a mild-temperate to humid subtropical lichen found in humid, old, open forests, on the trunks of old trees, occasionally on rocks and epilithic mosses. Most of the records are old; a recent record from Valle d'Aosta, a "continental" region, by Piervittori & Isocrono (1999) being dubious, is not accepted here. The species is included in the Italian red list of epiphytic lichens as “Near-threatened” (Nascimbene & al. 2013c).
Growth form: Foliose, broad lobed
Substrata: bark
Photobiont: cyanobacteria, filamentous (e.g. Nostoc, Scytonema)
Reproductive strategy: mainly asexual, by isidia, or isidia-like structures (e.g. schizidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: extremely rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: very rare
Humid mediterranean belt: extremely rare
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


Felix Schumm - CC BY-SA 4.0
[19062], Norfolk Island. Mt. Pitt Reserve, Selwyn Pine Track, 29°01' S, 167°56'30'' E, 200 m, on Elaedendron. Leg. J.A. Elix (18725) & H. Streimann, 08.12.1984. LICH. AUSTRALASICI EXS. 083



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5240)
2001/11/26
detail of isidia



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5240)
2001/11/26


Triin Aimla; Owner: University of Tartu



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (30534)
2008.03.10



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (16294)
2008.03.10



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (30534)
2008.03.10



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5240)
2008.03.10



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5240)
2008.03.10



E. Pittao; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (5240)
2008.03.10


Gothamie Weerakoon; Owner: Gothamie Weerakoon


Gothamie Weerakoon; Owner: Gothamie Weerakoon



Felix Schumm - CC BY-SA 4.0
[14524], Seychellen, La Digue, zwischen Belle Vue (Nid d' Aigles) und Fond Piment, 4°22' S, 55°50' E, ca. 350 m. Leg. F. Schumm & J.-P. Frahm 03.10.2008, det. A. Aptroot 2008.



Felix Schumm - CC BY-SA 4.0
[14524], Seychellen, La Digue, zwischen Belle Vue (Nid d' Aigles) und Fond Piment, 4°22' S, 55°50' E, ca. 350 m. Leg. F. Schumm & J.-P. Frahm 03.10.2008, det. A. Aptroot 2008.



Felix Schumm - CC BY-SA 4.0
[14524], Seychellen, La Digue, zwischen Belle Vue (Nid d' Aigles) und Fond Piment, 4°22' S, 55°50' E, ca. 350 m. Leg. F. Schumm & J.-P. Frahm 03.10.2008, det. A. Aptroot 2008.



Felix Schumm - CC BY-SA 4.0
[14524], Seychellen, La Digue, zwischen Belle Vue (Nid d' Aigles) und Fond Piment, 4°22' S, 55°50' E, ca. 350 m. Leg. F. Schumm & J.-P. Frahm 03.10.2008, det. A. Aptroot 2008.



Felix Schumm - CC BY-SA 4.0
[19062], Norfolk Island. Mt. Pitt Reserve, Selwyn Pine Track, 29°01' S, 167°56'30'' E, 200 m, on Elaedendron. Leg. J.A. Elix (18725) & H. Streimann, 08.12.1984. LICH. AUSTRALASICI EXS. 083



Felix Schumm - CC BY-SA 4.0
[19062], Norfolk Island. Mt. Pitt Reserve, Selwyn Pine Track, 29°01' S, 167°56'30'' E, 200 m, on Elaedendron. Leg. J.A. Elix (18725) & H. Streimann, 08.12.1984. LICH. AUSTRALASICI EXS. 083



Felix Schumm - CC BY-SA 4.0
[19062], Norfolk Island. Mt. Pitt Reserve, Selwyn Pine Track, 29°01' S, 167°56'30'' E, 200 m, on Elaedendron. Leg. J.A. Elix (18725) & H. Streimann, 08.12.1984. LICH. AUSTRALASICI EXS. 083



Felix Schumm - CC BY-SA 4.0
[19062], Norfolk Island. Mt. Pitt Reserve, Selwyn Pine Track, 29°01' S, 167°56'30'' E, 200 m, on Elaedendron. Leg. J.A. Elix (18725) & H. Streimann, 08.12.1984. LICH. AUSTRALASICI EXS. 083



Felix Schumm - CC BY-SA 4.0
[19062], Norfolk Island. Mt. Pitt Reserve, Selwyn Pine Track, 29°01' S, 167°56'30'' E, 200 m, on Elaedendron. Leg. J.A. Elix (18725) & H. Streimann, 08.12.1984. LICH. AUSTRALASICI EXS. 083



Felix Schumm - CC BY-SA 4.0
[19062], Norfolk Island. Mt. Pitt Reserve, Selwyn Pine Track, 29°01' S, 167°56'30'' E, 200 m, on Elaedendron. Leg. J.A. Elix (18725) & H. Streimann, 08.12.1984. LICH. AUSTRALASICI EXS. 083



Felix Schumm - CC BY-SA 4.0
[19062], Norfolk Island. Mt. Pitt Reserve, Selwyn Pine Track, 29°01' S, 167°56'30'' E, 200 m, on Elaedendron. Leg. J.A. Elix (18725) & H. Streimann, 08.12.1984. LICH. AUSTRALASICI EXS. 083



Juri Nascimbene
Altopiano del Canwsiglio, Regione Veneto, Prov. Treviso, Italy
2005



Felix Schumm – CC BY-SA 4.0
Image from: F. Schumm (2008) - Flechten Madeiras, der Kanaren und Azoren. Beck, OHG - ISBN: 978-3-00-023700-3



Curtis Randall Björk – CC BY-SA 4.0
British Columbia, Sabretooth Rapids, along the Clearwater River, Wells Gray Provincial Park Date: 2010-11-01 On rivershore rocks, in mistzone along whitewater


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Leif Stridvall - Source: http://www.stridvall.se/lichens/gallery/
Courtesy: Anita Stridvall


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.