Description: Thallus minutely squamulose, forming small aggregates several mm across. Squamules 0.1-0.5 mm wide, more or less crowded, adnate to ascending, flat to convex, 0.1-0.4 mm broad and to 0.5(-1) mm long, crenate to sublobate at margin, flattened to slightly convex, sometimes articulate in the form of a Opuntia, dispersed or clustered, rarely imbricate. Upper surface greyish brown or green-brown to fawn, green when wet, dull; lower surface pale. Thallus paraplectenchymatous throughout, with a poorly differentiated cortex, but upper cells minutely papillose, the algal cells distributed throughout most of the thallus. Perithecia prominent, arising between squamules, black, barrel-shaped, 0.2-0.6 mm broad, the surface rough and wrinkled. Exciple multi-layered, the outer part of densely pigmented, rounded cells, the middle part of unpigmented cells, the innermost layer of elongated, colourless cells; hamathecium of periphyses and periphysoids, interascal filaments absent; hymenial gel hemiamyloid I+ red, K/I+ blue. Asci (1-)2-spored, clavate, thin-walled, fissitunicate, not or very weakly thickened above, I-, K/I-. Ascospores strongly muriform, hyaline to pale yellow-brown, elongate-ellipsoid, (42-)80-120(-150) x (26-)30-50 μm. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a probably holarctic species with a wide altitudinal and latitudinal range, found on terricolous mosses, but also - albeit rarely - on basal parts of old trunks in calcareous areas.
Growth form: Squamulose
Substrata: soil, terricolous mosses, and plant debris