Cladonia ramulosa (With.) J.R. Laundon
Lichenologist, 16: 225, 1984. Basionym: Lichen ramulosus With. - Bot. Arrang. Veget. Gr. Brit.: 723, 1776.
Synonyms: Baeomyces anomaeus Ach.; Capitularia pityrea Flörke; Cladonia anomaea (Ach.) Ahti & P. James; Cladonia anomaea var. gracilior (Nyl.) Clauzade & Cl. Roux comb. inval.; Cladonia anomaea var. scyphifera (Delise) Clauzade & Cl. Roux comb. inval.; Cladonia anomaea var. subuliformis (Vain.) Clauzade & Cl. Roux comb. inval.; Cladonia degenerans var. anomaea (Ach.) Cromb.; Cladonia pityrea (Flörke) Fr.; Cladonia pseudopityrea Vain.
Distribution: N - Frl (Molaro 2005, Tretiach & Molaro 2007), Ven, TAA (Nascimbene & al. 2007b, 2022), Lomb (Rivellini & Valcuvia 1996, Gheza 2018, 2019b, Gheza & al. 2022), Piem (Isocrono & al. 2003, 2004), VA (Borlandelli & al. 1996, Piervittori & Isocrono 1997, Piervittori & al. 2001), Lig (Brunialti & al. 1999, Gheza & al. 2020). C - Tosc (Benesperi 2001, Ravera & al. 2019b, Gheza & al. 2020), Laz, Sar (Ahti & Puntillo 1995,Nöske 2000, Zedda & Sipman 2001, Zedda 2002). S - Camp (Ricciardi & al. 2000), Cal (Ahti & Puntillo 1995, Puntillo 1995, 1996), Si (Grillo & al. 1996, Grillo 1998, Czeczuga & al. 1999, Grillo & Caniglia 2004).
Description: Primary thallus squamulose, persistent or evanescent, the squamules small, 0.5-2(-5) mm long, 0.5-2 mm wide, elongate and deeply indented, fragile, glaucous green to greenish brown above, white beneath, rarely granulose-sorediate. Podetia simple or sparingly branched, hollow inside, greenish brown to brown, 0.5-3(-4) cm tall, 0.5-1(-3) mm thick, with pointed apices or with irregular, dentate and often marginally proliferating, small cups, squamulose-granular to partly decorticated, the areas without cortex usually with corticate granules or squamules, most abundant inside the cups. Apothecia frequent, pale to dark brown, convex, rather small, usually forming glomerules. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia developing on basal squamules and along margins of cups, subpedicellate, constricted at base, with a colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P+ red, UV-. Chemistry: fumarprotocetraric acid complex, occasionally with low amounts of atranorin and zeorin.Note: a mainly temperate to southern boreal-montane lichen found on epilithic bryophytes, rotting wood and organic soil, most common in upland areas with siliceous substrata. Specimens with delicate podetia with cortex plates detaching from the surface were described as C. pseudopityrea, which is distributed in temperate-humid climates of the Mediterranean region (Ahti & Puntillo 1995). However, according to Burgaz & al. (2020), unpublished molecular data do not support the hypothesis that C. pseudopityrea is different from C. ramulosa, and all European record of the N American species C. prolifica should be included in the variability of C. ramulosa, and European records of C. prolifica should be included in the variability of C. ramulosa.
Growth form: Fruticose
Substrata: bark, lignum, soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: rare
Oromediterranean belt: extremely rare
Montane belt: rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: rare
Humid mediterranean belt: rare
Dry mediterranean belt: absent
Predictive model
Herbarium samples
Image uploaded by Paul Cannon - CC BY-SA NC - Source: http://fungi.myspecies.info/all-fungi/cladonia-ramulosa
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10238)
2001/12/24
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Felix Schumm – CC BY-SA 4.0
[15959], Australia, Western Australia, north-east of Keysbrook, 32.41322° S, 116.01734° E, 260 m. Leg. F. Schumm & E. Stocker- Wörgötter, 29.10.2009, det. T. Ahti.
Felix Schumm – CC BY-SA 4.0
[15959], Australia, Western Australia, north-east of Keysbrook, 32.41322° S, 116.01734° E, 260 m. Leg. F. Schumm & E. Stocker- Wörgötter, 29.10.2009, det. T. Ahti.
Felix Schumm – CC BY-SA 4.0
[17489], Portugal, Azoren, Sao Miguel, zwischen Provoacao und Nordeste, Miradoro Tronqueira, Area Protegida da Tronqueira e planalto Graminhais, 37.79635° N, 25.18474° W, 682 m, Laurus-Erica Wald. Leg. Schumm & Frahm 24.09.2011, det. Schumm 2013
Felix Schumm – CC BY-SA 4.0
[17489], Portugal, Azoren, Sao Miguel, zwischen Provoacao und Nordeste, Miradoro Tronqueira, Area Protegida da Tronqueira e planalto Graminhais, 37.79635° N, 25.18474° W, 682 m, Laurus-Erica Wald. Leg. Schumm & Frahm 24.09.2011, det. Schumm 2013
Felix Schumm – CC BY-SA 4.0
[17489], Portugal, Azoren, Sao Miguel, zwischen Provoacao und Nordeste, Miradoro Tronqueira, Area Protegida da Tronqueira e planalto Graminhais, 37.79635° N, 25.18474° W, 682 m, Laurus-Erica Wald. Leg. Schumm & Frahm 24.09.2011, det. Schumm 2013
Felix Schumm – CC BY-SA 4.0
[15959], Australia, Western Australia, north-east of Keysbrook, 32.41322° S, 116.01734° E, 260 m. Leg. F. Schumm & E. Stocker- Wörgötter, 29.10.2009, det. T. Ahti.,
p: protocetraric acid, f: fumarprotocetraric acid, 2: physodalic acid, 1: homosekikaic acid?; conf. J.A. Elix
Felix Schumm – CC BY-SA 4.0
[19464], Germany, Hessen, Amöneburg bei Marburg, sonnige Basaltblöcke in einer Hangwiese kurz vor dem Ortseingang, 50.7896° N, 8.91995° E, 324 m. Leg. F. Schumm, 01.08.2016, det. F. Schumm.,
at: atranorin, ra: rangiformic acid, f: fumarprotocetraric acid (tr), p: protocetraric acid (tr)
Felix Schumm – CC BY-SA 4.0
[17489], Portugal, Azoren, Sao Miguel, zwischen Provoacao und Nordeste, Miradoro Tronqueira, Area Protegida da Tronqueira e planalto Graminhais, 37.79635° N, 25.18474° W, 682 m, Laurus-Erica Wald. Leg. Schumm & Frahm 24.09.2011, det. Schumm 2013
Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 550 m
03/2017
forms corresponding to C. pseudopityrea
Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 550 m
03/2017
forms corresponding to C. pseudopityrea
Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 550 m
03/2017
forms corresponding to C. pseudopityrea
Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 550 m
03/2017
forms corresponding to C. pseudopityrea
Giulio Pandeli
Italy
Giulio Pandeli
Italy
Giulio Pandeli
Italy
Giulio Pandeli
Italy
Giulio Pandeli
Italy
Giulio Pandeli
Italy
reaction to P
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Pointe de Rostudel
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Pointe de Treberon
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Argenton
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Argenton
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Argenton
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Briec
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Briec
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Briec
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Roscoff
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Canary Islands; La Gomera-MN; s of Las Rosas; s of Roque Blanco: near Mirador Dinero.
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Canary Islands; La Gomera-MN; s of Las Rosas; s of Roque Blanco: near Mirador Dinero.
Growth form: Fruticose
Substrata: bark, lignum, soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)
Most common in areas with a humid-warm climate (e.g. most of Tyrrenian Italy)
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: rare
Oromediterranean belt: extremely rare
Montane belt: rare
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: rare
Humid mediterranean belt: rare
Dry mediterranean belt: absent
Predictive model
| Herbarium samples |
Index Fungorum
GBIF
Image uploaded by Paul Cannon - CC BY-SA NC - Source: http://fungi.myspecies.info/all-fungi/cladonia-ramulosa
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (10238)
2001/12/24
Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php
Felix Schumm – CC BY-SA 4.0
[15959], Australia, Western Australia, north-east of Keysbrook, 32.41322° S, 116.01734° E, 260 m. Leg. F. Schumm & E. Stocker- Wörgötter, 29.10.2009, det. T. Ahti.
Felix Schumm – CC BY-SA 4.0
[15959], Australia, Western Australia, north-east of Keysbrook, 32.41322° S, 116.01734° E, 260 m. Leg. F. Schumm & E. Stocker- Wörgötter, 29.10.2009, det. T. Ahti.
Felix Schumm – CC BY-SA 4.0
[17489], Portugal, Azoren, Sao Miguel, zwischen Provoacao und Nordeste, Miradoro Tronqueira, Area Protegida da Tronqueira e planalto Graminhais, 37.79635° N, 25.18474° W, 682 m, Laurus-Erica Wald. Leg. Schumm & Frahm 24.09.2011, det. Schumm 2013
Felix Schumm – CC BY-SA 4.0
[17489], Portugal, Azoren, Sao Miguel, zwischen Provoacao und Nordeste, Miradoro Tronqueira, Area Protegida da Tronqueira e planalto Graminhais, 37.79635° N, 25.18474° W, 682 m, Laurus-Erica Wald. Leg. Schumm & Frahm 24.09.2011, det. Schumm 2013
Felix Schumm – CC BY-SA 4.0
[17489], Portugal, Azoren, Sao Miguel, zwischen Provoacao und Nordeste, Miradoro Tronqueira, Area Protegida da Tronqueira e planalto Graminhais, 37.79635° N, 25.18474° W, 682 m, Laurus-Erica Wald. Leg. Schumm & Frahm 24.09.2011, det. Schumm 2013
Felix Schumm – CC BY-SA 4.0
[15959], Australia, Western Australia, north-east of Keysbrook, 32.41322° S, 116.01734° E, 260 m. Leg. F. Schumm & E. Stocker- Wörgötter, 29.10.2009, det. T. Ahti.,
p: protocetraric acid, f: fumarprotocetraric acid, 2: physodalic acid, 1: homosekikaic acid?; conf. J.A. Elix
Felix Schumm – CC BY-SA 4.0
[19464], Germany, Hessen, Amöneburg bei Marburg, sonnige Basaltblöcke in einer Hangwiese kurz vor dem Ortseingang, 50.7896° N, 8.91995° E, 324 m. Leg. F. Schumm, 01.08.2016, det. F. Schumm.,
at: atranorin, ra: rangiformic acid, f: fumarprotocetraric acid (tr), p: protocetraric acid (tr)
Felix Schumm – CC BY-SA 4.0
[17489], Portugal, Azoren, Sao Miguel, zwischen Provoacao und Nordeste, Miradoro Tronqueira, Area Protegida da Tronqueira e planalto Graminhais, 37.79635° N, 25.18474° W, 682 m, Laurus-Erica Wald. Leg. Schumm & Frahm 24.09.2011, det. Schumm 2013
Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 550 m
03/2017
forms corresponding to C. pseudopityrea
Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 550 m
03/2017
forms corresponding to C. pseudopityrea
Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 550 m
03/2017
forms corresponding to C. pseudopityrea
Gabriele Gheza - https://lichenidilombardia.home.blog
Italy, Liguria, Imperia, Badalucco, 550 m
03/2017
forms corresponding to C. pseudopityrea
Giulio Pandeli
Italy
Giulio Pandeli
Italy
Giulio Pandeli
Italy
Giulio Pandeli
Italy
Giulio Pandeli
Italy
Giulio Pandeli
Italy
reaction to P
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Pointe de Rostudel
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Pointe de Treberon
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Argenton
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Argenton
Alain Gerault - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Argenton
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Briec
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Briec
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Briec
Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Roscoff
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=291&lang=en
France, Ardennes
Ulrich Kirschbaum CC BY-SA 4.0 - Source: https://www.thm.de/lse/ulrich-kirschbaum/flechtenbilder
Canary Islands; La Gomera-MN; s of Las Rosas; s of Roque Blanco: near Mirador Dinero.
