Cladonia subulata (L.) F.H. Wigg.

Cladonia subulata (L.) F.H. Wigg.

Primit. Florae Holsat.: 90, 1780. Basionym: Lichen subulatus L. - Sp. Pl.: 1153, 1753.

Synonyms: Cladonia cornutoradiata (Leight.) Sandst; Cladonia fimbriata var. cornutoradiata (Leight.) Vain.; Cladonia fimbriata var. radiata (Schreb.) Cromb.; Cladonia fimbriata var. subcornuta Nyl. ex Cromb.; Cladonia fimbriata var. subulata (L.) Vain.; Cladonia subulata var. radiata (Schreb.) Ozenda & Clauzade

Distribution: N - Frl (Tretiach & Molaro 2007), Ven (Caniglia & al. 1999, Nascimbene 2002, 2008, Nascimbene & Caniglia 2003c), TAA (Nascimbene & Caniglia 2000, Nascimbene 2006c, 2008b, Nascimbene & al. 2022), Lomb (Nascimbene 2006), Piem (Isocrono & al. 2004), Emil (Fariselli & al. 2020), Lig (Brunialti & al. 1999, Gheza & al. 2020). C - Tosc (Benesperi & al. 2007). S - Cal (Puntillo 1996), Si (Burgaz & al. 2019).

Description: Primary thallus squamulose, usually evanescent, the squamules very small, 1-2(-5) mm long, 2-3 mm wide, elongate and deeply incised, grey-green to whitish-glaucescent above, white beneath, sometimes sorediate at margins. Podetia simple or sparingly branched, the branchlets tending to diverge becoming antler-like, hollow inside, (2-)3-5(-10) cm tall, 0.3-1.5(-3) mm thick, esquamulose, with pointed apices, rarely with irregular small cups proliferating from margins, ash grey to whitish grey, sometimes with a greenish or brownish tinge, slightly melanotic at base, fully ecorticate or rarely corticate only at the very base, farinose-sorediate for most of their length, the soredia 20-50 μm in diam. Apothecia rare, terminal or along the margins of cups, brown, convex. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia rare, usually terminal on tips of podetia, with a colourless jelly. Conidia hyaline, curved. Photobiont chlorococcoid. Spot tests: K-, C-, KC-, P+ red, UV-. Chemistry: fumarprotocetraric acid complex.

Note: a cool-temperate to subarctic lichen found on mineral acidic soil on track sides and in clearings of light forests and heaths, more rarely on rotting wood or at the base of trunks, in upland areas. For the differences towards C. rei see Pino-Bodas & al. (2010b).

Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris

Photobiont: green algae other than Trentepohlia

Reproductive strategy: mainly asexual, by soredia, or soredia-like structures (e.g. blastidia)

Commonnes-rarity: (info)

Alpine belt: absent

Subalpine belt: very rare

Oromediterranean belt: extremely rare

Montane belt: very rare

Submediterranean belt: absent

Padanian area: absent

Humid submediterranean belt: absent

Humid mediterranean belt: absent

Dry mediterranean belt: absent

pH of the substrata:

1

2

3

4

5

(info)

Solar irradiation:

1

2

3

4

5

(info)

Aridity:

1

2

3

4

5

(info)

Eutrophication:

1

2

3

4

5

(info)

Poleotolerance:

0

1

2

3

(info)

Altitudinal distribution:

1

2

3

4

5

6

(info)

Predictive model

Herbarium samples

Leif Stridvall - Source: http://www.stridvall.se/la/galleries.php

Triin Aimla; Owner: University of Tartu

Felix Schumm – CC BY-SA 4.0

[8467], Portugal, Madeira, Rabacal, Lorbeerwald bei 25 Fontes, 32°45.699’ N, 17°8.051’ W, 1069 m. Leg. Schumm 12.04.2001, det. Aptroot 2010.

Felix Schumm – CC BY-SA 4.0

[8467], Portugal, Madeira, Rabacal, Lorbeerwald bei 25 Fontes, 32°45.699’ N, 17°8.051’ W, 1069 m. Leg. Schumm 12.04.2001, det. Aptroot 2010.

Felix Schumm – CC BY-SA 4.0

[8467], Portugal, Madeira, Rabacal, Lorbeerwald bei 25 Fontes, 32°45.699’ N, 17°8.051’ W, 1069 m. Leg. Schumm 12.04.2001, det. Aptroot 2010.

Felix Schumm – CC BY-SA 4.0

[8467], Portugal, Madeira, Rabacal, Lorbeerwald bei 25 Fontes, 32°45.699’ N, 17°8.051’ W, 1069 m. Leg. Schumm 12.04.2001, det. Aptroot 2010.

Felix Schumm – CC BY-SA 4.0

[17286], Germany, Baden-Württemberg, Rems-Murr-Kreis, Stetten im Remstal, Haldenbachtal, Wegrand im Nadelwald. Leg. Erich Putzler 05.04.1950, det. J.Poelt.

Felix Schumm – CC BY-SA 4.0

[19490], Germany, Schwarzwald, Kreis Freudenstadt, NW von Baiersbronn, ca. 2 km oberhalb Obertal. 48.54692° N, 8.27071° E, 720 m. Leg. F. Schumm, 6.9.2016, det. F. Schumm

Felix Schumm – CC BY-SA 4.0

[19490], Germany, Schwarzwald, Kreis Freudenstadt, NW von Baiersbronn, ca. 2 km oberhalb Obertal. 48.54692° N, 8.27071° E, 720 m. Leg. F. Schumm, 6.9.2016, det. F. Schumm

Felix Schumm – CC BY-SA 4.0

[19490], Germany, Schwarzwald, Kreis Freudenstadt, NW von Baiersbronn, ca. 2 km oberhalb Obertal. 48.54692° N, 8.27071° E, 720 m. Leg. F. Schumm, 6.9.2016, det. F. Schumm

Felix Schumm – CC BY-SA 4.0

[8467], Portugal, Madeira, Rabacal, Lorbeerwald bei 25 Fontes, 32°45.699’ N, 17°8.051’ W, 1069 m. Leg. Schumm 12.04.2001, det. Aptroot 2010.,
HPTLC in den Laufmitteln A, B’ und C nach Säurebehandlung bei Tageslicht und 366 nm UVLangwelle. 1: Start, 4: Position von Norstictinsäure 7: Position von Atranorin pr: Protocetrarsäure fpr: Fumarprotocetrarsäure

Curtis Randall Björk – CC BY-SA 4.0

British Columbia, Clearwater Valley 2014-04-00 On rock in waterfall sprayzone

Leif Stridvall

Courtesy: Anita Stridvall -Source: http://www.stridvall.se/lichens/gallery/

Mark Rahill - CC BY-SA 4.0

USA

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Col d'Iraty

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Col d'Iraty

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Col d'Iraty

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Commana

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Commana

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Ardennes

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Ardennes

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Ardennes

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Ardennes

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=561&lang=en

France, Ardennes

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=392&lang=en

France, Cap de la Chèvre
var. radiata

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=392&lang=en

France, Cap de la Chèvre
var. radiata

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=392&lang=en

France, Cap de la Chèvre
var. radiata

Bernard Bouffinier - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=392&lang=en

France, Cap de la Chèvre
var. radiata

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