Cladonia uncialis (L.) F.H. Wigg. subsp. biuncialis (Hoffm.) M. Choisy

Bull. Mens. Soc. Linn. Lyon, 20: 9, 1951. Basionym: Cladonia biuncialis Hoffm. - Deutschl. Fl., 2: 116, 1796.
Synonyms: Cladonia uncialis f. turgescens (Delise) Fr.; Cladonia uncialis subsp. dicraea (Ach.) D. Hawksw.
Distribution: N - Frl (Martellos 2005), Ven (Jatta 1909-1911).
Description: Primary thallus squamulose, ephemeral and rarely seen, the squamules to 1 mm long, white beneath. Podetia shrubby, forming more or less compact tufts, hollow inside, pale yellowish green, pale grey and not melanotic at base, the terminal branchlets often brownish at tips, esquamulose, sparingly and mainly anisotomic-dichotomously branched, to 2-5(-8) cm tall, 0.5-1 mm thick, with pointed apices and divergent branches, the axils perforate or closed. Cortex smooth, maculate; surface of central canal finely powdery-granulose, especially in upper parts. Apothecia extremely rare, brown, convex, small. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed, apical, with a colourless jelly. Conidia cylindrical. Photobiont chlorococcoid. Spot tests: K-, C-, KC+ yellow, P-, UV+ white (medulla). Chemistry: usnic and sometimes squamatic acids.
Note: this subspecies differs from subsp. uncialis in the frequent presence of squamatic acid, and in having perforated, (di-), tri- or tetrachotomous ultimate ramifications. Mixed population may occur, where ìndividuals may be difficult to assign to either subspecies.
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: extremely rare
Subalpine belt: rare
Oromediterranean belt: absent
Montane belt: absent
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:


Solar irradiation:








Altitudinal distribution:


Predictive model
Herbarium samples

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: GZU Herb.Hafellner (49998)

Harrie Sipman – Source - As Caloplaca oasis

Curtis Randall Björk – CC BY-SA 4.0
British Columbia, Skeena Mountains 2014-08-01 On gravel bed in alpine grassland

Pier Luigi Nimis CC BY-SA 4.0
TSB 38582

Pier Luigi Nimis CC BY-SA 4.0
TSB 38850

Pier Luigi Nimis CC BY-SA 4.0
TSB 387850

Courtesy Danièle et Olivier Gonnet - Source:
France, Session AFL 2009 - Fontainebleau

Courtesy Danièle et Olivier Gonnet - Source:
France, Session AFL 2009 - Fontainebleau