Cladonia uncialis (L.) F.H. Wigg. subsp. uncialis

Primit. Florae Holsat.: 90, 1780. Basionym: Lichen uncialis L. - Sp. Pl.: 1153, 1753, nom. cons.
Synonyms: Cladonia uncialis var. obtusata (Ach.) Räsänen
Distribution: N - Frl (Tretiach & Hafellner 2000), Ven (Caniglia & al. 1999, Nascimbene & Caniglia 2003c), TAA (Caniglia & al. 2002, Nascimbene & al. 2005, 2006, 2022, Brackel 2013, Bilovitz & al. 2014), Lomb (Rivellini 1994, Rivellini & Valcuvia 1996, Dalle Vedove & al. 2004, Gheza 2018, 2019, 2019b, Gheza & al. 2022), Piem (Morisi & Sereno 1995, Isocrono & al. 2004, 2006, Isocrono & Piervittori 2008, Gheza 2015, 2018), VA (Piervittori & Isocrono 1999), Emil (Dalle Vedove & al. 2002, Benesperi & al. 2007, Fariselli & al. 2020), Lig (Watson 2014). C - Tosc, Laz (Jatta 1909-1911), Sar.
Description: Primary thallus squamulose, ephemeral and rarely seen, the squamules to 1 mm long, white below. Podetia forming compact tufts, hollow inside, pale yellowish green, paler towards the base, often with brownish tips, esquamulose, mostly tri- to tetrachotomously branched, up to 6(-10) cm tall, c. 1.5(-2) mm thick, subcylindrical in section, cupless, with pointed apices and divergent terminal branchlets, the axils mostly perforate. Cortex smooth, maculate; surface of central canal not powdery. Apothecia rare, brown, convex, up to 2 mm across. Asci 8-spored, clavate, thickened at apex, with a K/I+ blue tholus and a K/I+ strongly blue outer gelatinous sheath, Cladonia-type. Ascospores 1-celled, hyaline, ellipsoid. Pycnidia dark, semi-immersed, apical, with a colourless jelly. Conidia cylindrical. Photobiont chlorococcoid. Spot tests: K-, C-, KC+ yellow, P-, UV- (rarely UV+ white). Chemistry: usnic acid, rarely also squamatic acid.
Note: an arctic-alpine to northern boreal-montane, circumpolar species found on soil and amongst mosses in very open habitats with a long snow-lie near or above treeline, common only in the Alps. In the western part of the upper Padanian Plain there are some dealpine stations in the submediterranean belt. Some records could refer to subsp. biuncialis.
Growth form: Fruticose

Substrata: soil, terricolous mosses, and plant debris
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: rare
Subalpine belt: rather rare
Oromediterranean belt: extremely rare
Montane belt: absent
Submediterranean belt: extremely rare
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:


Solar irradiation:








Altitudinal distribution:


Predictive model
Herbarium samples

Leif Stridvall - Source:

P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (1701)
tips of podetia

Triin Aimla; Owner: University of Tartu
originally as Cladonia uncialis

Gabriele Gheza; Owner: Gabriele Gheza
Italy, Lombardia, Alta Valtellina; altitude: c. 2000 m
summer 2010

Gabriele Gheza; Owner: Gabriele Gheza
Italy, Lombardia, Alta Valtellina; altitude: c. 2000 m
summer 2010

Juri Nascimbene – CC BY-SA 4.0; Owner: Department of ùLife Sciences, University of Trieste
Calaita, Italy

Mark Rahill - CC BY-SA 4.0
USA, Vermont

Pier Luigi Nimis - CC BY-SA 4.0
TSB 43966