Dermatocarpon intestiniforme (Körb.) Hasse

Bryologist, 15: 46, 1912. Basionym: Endocarpon intestiniforme Körb. - Parerga Lichenol.: 42, 1859.
Synonyms: Dermatocarpon aquaticum var. decipiens (A. Massal.) Zahlbr. non auct.; Dermatocarpon decipiens (A. Massal.) Dalla Torre & Sarnth. non auct.; Dermatocarpon fluviatile var. decipiens (A. Massal.) Vain. non auct.; Dermatocarpon luridum var. decipiens (A. Massal.) H. Riedl non auct.; Dermatocarpon polyphyllum Dalla Torre & Sarnth.; Dermatocarpon weberi var. decipiens (A. Massal.) Lambinon non auct.; Endocarpon decipiens A. Massal. non auct.; Endocarpon miniatum var. decipiens A. Massal. non auct.
Distribution: N - Frl, Ven (Nascimbene & Caniglia 2000, 2003c), TAA (Nascimbene & al. 2022), Lomb (Gheza 2019), Piem (Morisi & Sereno 1995, Isocrono & al. 2003, Morisi 2005), VA (Piervittori & Isocrono 1997, 1999). C - Tosc (TSB 36835), Umb (Ravera & Di Toma 2003, Ravera & al. 2006), Mol (Ravera & Genovesi 2012, Genovesi & Ravera 2014). S - Cal (Puntillo 1996).
Description: Thallus foliose to subcrustose, heteromerous, dorsiventral, not umbilicate, multi-lobed, forming up to 7(-10) cm wide patches, the lobes interconnected, with numerous holdfasts scattered over the lower surface, (3-) 5-15(-20) mm wide, the central ones markedly sinuous, with down-turned margins. Upper surface grey-brown, usually pruinose; lower surface brown to dark brown, smooth to rugose. Upper cortex (40-)50-90(-110) μm thick, pseudoparenchymatous, of anticlinallly arranged hyphae, with a brown pigment near the surface, overlain by an epinecral layer of air-filled hyphae; medulla white, of filamentous hyphae, I-; lower cortex of anticlinally arranged hyphae. Perithecia common, slightly oblong, immersed in the thallus and scattered over almost the whole upper surface (appearing as black dots), without involucrellum. Exciple colourless except in the brown uppermost part; hamathecium or periphyses and periphysoids, interascal filaments absent; hymenial gel hemiamyloid. Asci 8-spored, clavate, I-, fissitunicate, the wall thickened above, with an ocular chamber, dehiscent by extrusion of an endotunica to form a delicate rostrum, Verrucaria-type. Ascospores 1-celled, hyaline, broadly ellipsoid to subglobose, (8-)9-15(-16) x (5-)6-8(-9) µm. Pycnidia common, black, immersed. Conidia bacilliform. Photobiont chlorococcoid. Spot tests: cortex and medulla K-, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Note: a mainly boreal-montane to arctic-alpine, circumpolar lichen found on more or less calciferous rocks in periodically wet places, or near the ground, mostly in upland areas. According to Heidmarsson (2017), however, this is just a morph of D. miniatum. As the molecular taxonomy of the D. miniatum-complex is not fully resolved, I prefer to provisionally maintain the taxon as distinct.
Growth form: Foliose, narrow lobed
Substrata: rocks
Photobiont: green algae other than Trentepohlia
Reproductive strategy: mainly sexual
Periodically submerged (e.g. in creeks)

Commonnes-rarity: (info)

Alpine belt: rare
Subalpine belt: rather rare
Oromediterranean belt: extremely rare
Montane belt: extremely rare
Submediterranean belt: absent
Padanian area: absent
Humid submediterranean belt: absent
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples

Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Queyras



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35223)
2002/07/12



Curtis Randall Björk – CC BY-SA 4.0
Potlatch River Canyon, Latah County, Idaho Date: 2011-10-08 On river boulders at shoreline



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (35223)
2002/07/12
lower face


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_D/Dermatocarpon_intestiniforme.htm
France, session AFL 2014 Mercantour


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_D/Dermatocarpon_intestiniforme.htm
France, session AFL 2014 Mercantour


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_D/Dermatocarpon_intestiniforme.htm
France, session AFL 2014 Mercantour


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Beaufortin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Beaufortin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Beaufortin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Queyras



P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (34181)
2001/12/13


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Beaufortin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Beaufortin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Beaufortin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Beaufortin


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=969&lang=en
France, Beaufortin