Anema tumidulum P.M. Jørg., M. Schultz & Guttová

Henssen ex P.M. Jørg., M. Schultz & Guttová, Herzogia, 26: 2, 2013.
Synonyms:
Distribution: N - Frl, Ven (Nascimbene & al. 2021), Lomb (Jørgensen & al. 2013), VA (Matteucci & al. 2013). C - Tosc (Tretiach & al. 2008), Abr (Caporale & al. 2016). S - Cal (CLU 18152).
Description: Thallus squamulose, homoiomerous, slightly gelatinous when wet, in section with a loosely reticulate network of thin hyphae enclosing groups of cyanobacterial cells. Squamules shield-like, attached by a central point, up to 3(-4) mm wide, dispersed to usually crowded and forming effuse, black-brown crusts, with an often lobulate to incised margin, epruinose, often shiny, covered in granular isidia-like outgrowths which aggregate forming a dense bushy structure around the apothecia. Apothecia pycnoascocarps, at first immersed and perithecioid, but soon expanded and lecanorine, 0.5-0.8(-1) mm across, with a dark reddish brown disc and a thick thalline margin. Proper exciple thin, cupulate; epithecium yellowish brown; hymenium and hypothecium colourless, the hymenial gelatine I+ blue. Asci 4(-8)-spored, subcylindrical, thin-walled, with an amyloid external apical cap. Ascospores 1-celled, hyaline, broadly ellipsoid to subglobose, 8-13(-17) x (4-)6-8(-11) µm, the wall 1-2 µm thick. Pycnidia laminal, immersed. Conidia narrowly ellipsoid to bacilliform, c. 3 x 1 µm. Photobiont cyanobacterial, chroococcoid, of a few 4-10 µm wide cells surrounded by a gelatinous sheath which is colourless in inner parts, yellowish brown near the thallus surface. Spot tests: all negative. Chemistry: without lichen substances.
Note: this species seems to be fairly common in Central Europe (Jørgensen & al. 2013). It grows on steeply inclined, sunny surfaces of calcareous or basic siliceous rocks with periodical water seepage after rain, with optimum in upland areas. See also note on A. moedligense.
Growth form: Crustose
Substrata: rocks
Photobiont: cyanobacteria, coccaceous (e.g. Gloeocapsa)
Reproductive strategy: mainly sexual
On otherwise dry surfaces with short periods of water seepage after rain

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Oromediterranean belt: absent
Montane belt: rather rare
Submediterranean belt: rare
Padanian area: absent
Humid submediterranean belt: extremely rare
Humid mediterranean belt: absent
Dry mediterranean belt: absent

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Predictive model
Herbarium samples


P.L. Nimis CC BY-SA 04
TSB 34942


courtesy Olivier & Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A2/Anema_tumidulum.htm
France, Session AFL dans le Lot - Saint-Sozy
25/5/2015


courtesy Olivier & Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A2/Anema_tumidulum.htm
France, Session AFL dans le Lot - Saint-Sozy
25/5/2015


courtesy Olivier & Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A2/Anema_tumidulum.htm
France, Session AFL dans le Lot - Saint-Sozy
25/5/2015


courtesy Olivier & Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A2/Anema_tumidulum.htm
France, Session AFL dans le Lot - Saint-Sozy
25/5/2015


courtesy Olivier & Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A2/Anema_tumidulum.htm
France, Session AFL dans le Lot - Saint-Sozy
25/5/2015


courtesy Olivier & Danièle Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_A/Textes_A2/Anema_tumidulum.htm
France, Session AFL dans le Lot - Saint-Sozy
25/5/2015