Graphis betulina (Pers.) Ach.
K. Vetensk.-Akad. Nya Handl., 30: 147, 1809. Basionym: Opegrapha betulina Pers. - Ann. Bot. (Usteri), 7: 31, 1794.
Synonyms: Graphis juglandis Garov. ex A. Massal.; Graphis massalongii Kremp.
Description: Thallus crustose, continuous to finely cracked, white, dull, often delimited by a dark prothallus. Apothecia lirelliform, black, raised or embedded in the thallus, oblong to linear, more or less flexuose, branched and acute at the ends, 2-8 x 0.1-0.3 mm, with a slit-like or at least centrally expanded, pruinose or epruinose disc, a well-developed, 15-45 µm thick proper margin with entire lips, and a conspicuous, c. 0.2 mm thick, pseudothalline margin filled with calcium oxalate crystals. Proper exciple carbonized laterally, poorly developed and not carbonized at base; epithecium brown; hymenium colourless, not inspersed with oil droplets, 90-100 µm high; paraphyses simple, dense, 1.5-2 µm thick, with brownish tips; subhymenium colourless. Asci (6-)8-spored, cylindrical-clavate, non fissitunicate, dehiscent by an apical split, K/I-, Graphis-type. Ascospores 5-11-septate with lens-shaped lumina, hyaline (pale brown when overmature), 15-45 x 5-9 µm, the wall I+ blue-violet. Pycnidia rare, immersed. Conidia bacilliform, 2-5 x 1 µm. Photobiont trentepohlioid. Spot tests: cortex and medulla K- or K+ yellowish brown, C-, KC-, P-. Chemistry: without lichen substances. 
					Growth form: Crustose
Substrata: bark
Photobiont: Trentepohlia
Reproductive strategy: mainly sexual
		Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Montane belt: rare
Dry submediterranean belt: rare
Humid submediterranean belt: rather rare
Padanian area: absent
	
					
        
          
            pH of the substrata:
          
          
            
              
                1 2 3 4 5 
            
          
          
          
        
          
        
          
					
        
          
            Solar irradiation:
          
          
            
              
                1 2 3 4 5 
            
          
          
          
        
          
        
          
					
        
          
            Aridity:
          
          
            
              
                1 2 3 4 5 
            
          
          
          
        
          
        
          
					
        
          
            Eutrophication:
          
          
            
              
                1 2 3 4 5 
            
          
          
          
        
          
        
          
					
        
          
            Poleotolerance:
          
          
            
              
                0 1 2 3 
            
          
          
          
        
          
        
          
					
        
          
            Altitudinal distribution:
          
          
            
              
                1 2 3 4 5 6 
            
          
          
          
        
          
        
           
    	  
        
        
    Rarity
    
    
      
      absent
    
    
    
      
      extremely rare
    
    
    
      
      very rare
    
    
    
      
      rare
    
    
    
      
      rather rare
    
    
    
      
      rather common
    
    
    
      
      common
    
    
    
      
      very common
    
    
    
      
      extremely common
    
  
			
      
			
    	  
        Loading data...
			
			
      
			
        Occurrence data
        
			
			
      
			
        Predictive map
       
			
			
			
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_betulina.htm
France, session AFL 2015 - sur Corylus - La Chapelle Auzac - Blagour - Lot

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_betulina.htm
France, session AFL 2015 - sur Corylus - La Chapelle Auzac - Blagour - Lot

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_betulina.htm
France, session AFL 2015 - sur Corylus - La Chapelle Auzac - Blagour - Lot
        
        
        
        
        
        
        
        
        
        
        
        
        
        
        
    
				
			
                        
Growth form: Crustose
Substrata: bark
Photobiont: Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Montane belt: rare
Dry submediterranean belt: rare
Humid submediterranean belt: rather rare
Padanian area: absent
pH of the substrata:
| 1 | 2 | 3 | 4 | 5 | 
Solar irradiation:
| 1 | 2 | 3 | 4 | 5 | 
Aridity:
| 1 | 2 | 3 | 4 | 5 | 
Eutrophication:
| 1 | 2 | 3 | 4 | 5 | 
Poleotolerance:
| 0 | 1 | 2 | 3 | 
Altitudinal distribution:
| 1 | 2 | 3 | 4 | 5 | 6 | 
Rarity
      
      absent
    
    
    
      
      extremely rare
    
    
    
      
      very rare
    
    
    
      
      rare
    
    
    
      
      rather rare
    
    
    
      
      rather common
    
    
    
      
      common
    
    
    
      
      very common
    
    
    
      
      extremely common
    
  Loading data...
Occurrence data
        
			Predictive map
       
			
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_betulina.htm
France, session AFL 2015 - sur Corylus - La Chapelle Auzac - Blagour - Lot

Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_betulina.htm
France, session AFL 2015 - sur Corylus - La Chapelle Auzac - Blagour - Lot







 
                                              
                                              
                                              
                                             