Graphis scripta (L.) Ach.

Lichenogr. Univ.: 265, 1810. Basionym: Lichen scriptus L. - Sp. Pl.: 1140, 1753.
Synonyms: Graphis hebraica (Hoffm.) Röhl.; Graphis limitata (Pers.) Röhl.; Graphis microcarpa (Ach.) Röhl.; Graphis scripta var. limitata (Pers.) Arnold; Graphis scripta var. spathea (Ach.) Mudd; Graphis spathea (Ach.) Röhl.
Description: Thallus crustose, continuous to finely cracked, white, pale grey or brownish, dull, often delimited by a dark prothallus. Apothecia lirelliform, black, raised, oblong to linear, more or less flexuose and often with several parallel branches, rounded at ends, 2-10 x 0.2-0.4 mm, with a slit-like, epruinose disc hidden by a well-developed, often glossy, 15-40 µm thick exciple with entire lips; a very thin pseudothalline margin is often present. Proper exciple black, carbonized laterally, poorly developed and not carbonized at base; epithecium brown; hymenium colourless, not inspersed with oil droplets, 90-100 µm high; paraphyses simple, dense, 1.5-2 µm thick, with brownish tips; subhymenium colourless. Asci (6-)8-spored, clavate or cylindrical-clavate, non fissitunicate, dehiscent by an apical split, K/I-, Graphis-type. Ascospores: (5-)7-10(-13)-septate with lens-shaped lumina, hyaline, fusiform-cylindrical, 20-63 x 5-12 µm, the wall I+ blue-violet. Pycnidia rare, immersed. Conidia bacilliform, 2-5 x 1 µm. Photobiont trentepohlioid. Spot tests: cortex and medulla K- or K+ yellowish brown, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Growth form: Crustose
Substrata: bark
Photobiont: Trentepohlia
Reproductive strategy: mainly sexual

Commonnes-rarity: (info)

Alpine belt: absent
Subalpine belt: absent
Montane belt: very common
Dry submediterranean belt: rare
Humid submediterranean belt: rather rare
Padanian area: extremely rare

pH of the substrata:

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Solar irradiation:

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Aridity:

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Eutrophication:

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Poleotolerance:

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Altitudinal distribution:

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Rarity

absent
extremely rare
very rare
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common
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extremely common
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Occurrence data
Predictive map
Model ImageCurrent prediction (1981-2010)
Model ImageFuture prediction (2071-2100) SSP 1-2.6
Model ImageFuture prediction (2071-2100) SSP 5-8.5
Predictive maps according to Francesconi et al. 2025

G. Incerti CC BY-SA 4.0
TSB 31894


G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31894)
2001/11/26


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (17705)
2001/11/24


Andres Saag; Owner: University of Tartu


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08


Domenico Puntillo; Owner: Domenico Puntillo
Italy


P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Bosco della Stua, Lago di Sauris, 1000m
09.09.2016


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017


Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.


Juri Nascimbene
Altopiano del Canwsiglio, Regione Veneto, Prov. Treviso, Italy
2005


G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31894)
2001/11/26


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_scripta.htm
France, sur écorce de bouleau - Forêt d'Hardelot - 2007 - Pas-de-Calais


Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_scripta.htm
France, sur écorce de bouleau - Forêt d'Hardelot - 2007 - Pas-de-Calais


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France


Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Kempten


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Kempten


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Taubenberg


Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Taubenberg


Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.


Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italy, Veneto, Ubique in regno Veneto 1855