Graphis scripta (L.) Ach.
Lichenogr. Univ.: 265, 1810. Basionym: Lichen scriptus L. - Sp. Pl.: 1140, 1753.
Synonyms: Graphis hebraica (Hoffm.) Röhl.; Graphis limitata (Pers.) Röhl.; Graphis microcarpa (Ach.) Röhl.; Graphis scripta var. limitata (Pers.) Arnold; Graphis scripta var. spathea (Ach.) Mudd; Graphis spathea (Ach.) Röhl.
Description: Thallus crustose, continuous to finely cracked, white, pale grey or brownish, dull, often delimited by a dark prothallus. Apothecia lirelliform, black, raised, oblong to linear, more or less flexuose and often with several parallel branches, rounded at ends, 2-10 x 0.2-0.4 mm, with a slit-like, epruinose disc hidden by a well-developed, often glossy, 15-40 µm thick exciple with entire lips; a very thin pseudothalline margin is often present. Proper exciple black, carbonized laterally, poorly developed and not carbonized at base; epithecium brown; hymenium colourless, not inspersed with oil droplets, 90-100 µm high; paraphyses simple, dense, 1.5-2 µm thick, with brownish tips; subhymenium colourless. Asci (6-)8-spored, clavate or cylindrical-clavate, non fissitunicate, dehiscent by an apical split, K/I-, Graphis-type. Ascospores: (5-)7-10(-13)-septate with lens-shaped lumina, hyaline, fusiform-cylindrical, 20-63 x 5-12 µm, the wall I+ blue-violet. Pycnidia rare, immersed. Conidia bacilliform, 2-5 x 1 µm. Photobiont trentepohlioid. Spot tests: cortex and medulla K- or K+ yellowish brown, C-, KC-, P-, UV-. Chemistry: without lichen substances.
Growth form: Crustose
Substrata: bark
Photobiont: Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Montane belt: very common
Dry submediterranean belt: rare
Humid submediterranean belt: rather rare
Padanian area: extremely rare
pH of the substrata:
1 2 3 4 5
Solar irradiation:
1 2 3 4 5
Aridity:
1 2 3 4 5
Eutrophication:
1 2 3 4 5
Poleotolerance:
0 1 2 3
Altitudinal distribution:
1 2 3 4 5 6
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Current prediction (1981-2010)
Future prediction (2071-2100) SSP 1-2.6
Future prediction (2071-2100) SSP 5-8.5Predictive maps according to Francesconi et al. 2025
G. Incerti CC BY-SA 4.0
TSB 31894
G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31894)
2001/11/26
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (17705)
2001/11/24
Andres Saag; Owner: University of Tartu
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08
Domenico Puntillo; Owner: Domenico Puntillo
Italy
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Bosco della Stua, Lago di Sauris, 1000m
09.09.2016
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Juri Nascimbene
Altopiano del Canwsiglio, Regione Veneto, Prov. Treviso, Italy
2005
G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31894)
2001/11/26
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_scripta.htm
France, sur écorce de bouleau - Forêt d'Hardelot - 2007 - Pas-de-Calais
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_scripta.htm
France, sur écorce de bouleau - Forêt d'Hardelot - 2007 - Pas-de-Calais
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Kempten
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Kempten
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Taubenberg
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Taubenberg
Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.
Collezione lichenologica Abramo Massalongo del Museo di Storia Naturale G. Ligabue di Venezia - Autori: Seggi, Linda; Trabucco, Raffaella Proprietà: Fondazione Musei Civici di Venezia - CC BY-NC
Italy, Veneto, Ubique in regno Veneto 1855
Growth form: Crustose
Substrata: bark
Photobiont: Trentepohlia
Reproductive strategy: mainly sexual
Commonnes-rarity: (info)
Alpine belt: absent
Subalpine belt: absent
Montane belt: very common
Dry submediterranean belt: rare
Humid submediterranean belt: rather rare
Padanian area: extremely rare
pH of the substrata:
| 1 | 2 | 3 | 4 | 5 |
Solar irradiation:
| 1 | 2 | 3 | 4 | 5 |
Aridity:
| 1 | 2 | 3 | 4 | 5 |
Eutrophication:
| 1 | 2 | 3 | 4 | 5 |
Poleotolerance:
| 0 | 1 | 2 | 3 |
Altitudinal distribution:
| 1 | 2 | 3 | 4 | 5 | 6 |
Rarity
absent
extremely rare
very rare
rare
rather rare
rather common
common
very common
extremely common
Loading data...
Occurrence data
Predictive map
Current prediction (1981-2010)Future prediction (2071-2100) SSP 1-2.6Future prediction (2071-2100) SSP 5-8.5Predictive maps according to Francesconi et al. 2025
G. Incerti CC BY-SA 4.0
TSB 31894
G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31894)
2001/11/26
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (17705)
2001/11/24
Andres Saag; Owner: University of Tartu
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italia, Friuli-Venezia Giulia, UD, Comune di Sauris, Bosco della Stua presso il Lago
2008.26.08
Domenico Puntillo; Owner: Domenico Puntillo
Italy
P.L. Nimis; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Bosco della Stua, Lago di Sauris, 1000m
09.09.2016
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017
Andrea Moro; Owner: Department of Life Sciences, University of Trieste
Italy, Friuli Venezia Giulia, Udine, Lago di Sauris, Bosco della Stua, Carnic Alps (Ampezzo Carnico) 1000 m; altitude: 1000 m
12/08/2017
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Felix Schumm – CC BY-SA 4.0
[12550], Germany, Baden-Württemberg, Kreis Göppingen, zwischen Wangen und Unterberken, im Fetzentöbele, Weg ‘Seesteige’, an schattiger junger Buche, 48°45.494'N, 9°34.628'E, 390 m, TK 7223. Leg. et det. F. Schumm, 16.07.2007.
Juri Nascimbene
Altopiano del Canwsiglio, Regione Veneto, Prov. Treviso, Italy
2005
G. Incerti; Owner: Department of Life Sciences, University of Trieste
Herbarium: TSB (31894)
2001/11/26
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_scripta.htm
France, sur écorce de bouleau - Forêt d'Hardelot - 2007 - Pas-de-Calais
Courtesy Danièle et Olivier Gonnet - Source: https://www.afl-lichenologie.fr/Photos_AFL/Photos_AFL_G/Graphis_scripta.htm
France, sur écorce de bouleau - Forêt d'Hardelot - 2007 - Pas-de-Calais
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Jacques Haine - Source: http://www.lichensmaritimes.org/index.php?task=fiche&lichen=588&lang=en
France
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Kempten
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Kempten
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Taubenberg
Wolfgang von Brackel - CC BY-4.0
Germany, Bavaria, Taubenberg
Zahlbruckner A. 1926. Lichenes (Flechten). In: Engler A. (ed.): Die natürlichen Pflanzenfamilien. 2nd ed., vol 8, W. Engelmann, Leipzig, 270 pp.
